large beardtongue, largeflower penstemon, shell-leaf beardtongue
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Utah beardtongue, Utah penstemon
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erect, (40–)50–95(–120) cm, glabrous. |
ascending to erect, 15–60 cm, glaucous. |
basal and cauline, glabrous; basal and proximal cauline 30–160 × 6–50 mm, blade spatulate to obovate, base tapered, apex rounded to obtuse or acute; cauline 4–8 pairs, sessile, 18–90(–110) × 15–50 mm, blade spatulate to orbiculate, base clasping, apex rounded to obtuse. |
glabrous or obscurely scabrous, especially along margins, glaucous; basal and proximal cauline 35–80(–100) × 5–20 mm, blade oblanceolate, base tapered, margins entire, apex obtuse to acute; cauline 2–4 pairs, sessile, 15–75 × 4–26 mm, blade elliptic to lanceolate, rarely ovate, base tapered, margins entire, apex acute. |
interrupted, cylindric, 12–30(–40) cm, axis glabrous, verticillasters 3–7(–9), cymes 2–4-flowered; proximal bracts ovate to elliptic or orbiculate, (9–)16–83 × (9–)16–54 mm; peduncles and pedicels glabrous. |
± interrupted, ± secund, 11–25 cm, axis glabrous, verticillasters 5–15, cymes 1–4-flowered; proximal bracts lanceolate, 6–23 × 1–4 mm; peduncles and pedicels erect, glabrous. |
calyx lobes ovate to lanceolate, 7–11 × 2.5–4 mm, margins entire, rarely erose, herbaceous or narrowly scarious, glabrous; corolla lavender to blue or pinkish blue, with magenta nectar guides, ampliate, 35–48 mm, glabrous externally, glabrous internally, tube 10–13 mm, throat abruptly inflated, 15–18 mm diam., rounded abaxially; stamens included, pollen sacs opposite, 2.1–2.6 mm, sutures papillate; staminode 16–21 mm, included or reaching orifice, 2–2.6 mm diam., tip recurved to coiled, distal 1–2 mm sparsely villous, hairs golden yellow, to 0.5 mm; style 19–30 mm. |
calyx lobes ovate, 2.5–4(–5) × 1.5–4(–4.6) mm, margins erose, glabrous; corolla red to crimson, with red nectar guides, bilaterally symmetric, bilabiate, tubular-salverform, 17–22(–25) mm, glandular-pubescent externally, especially distally, glandular-pubescent internally, tube 6–9 mm, throat slightly inflated, 6–7 mm diam., rounded abaxially; stamens included, pollen sacs explanate, 0.6–1.1 mm, sutures smooth; staminode 7–10 mm, flattened distally, 0.4–0.5 mm diam., tip recurved, glabrous, rarely distal 1 mm papillate, papillae golden yellow, to 0.1 mm; style 10–14 mm. |
16–20(–25) × 8–15 mm. |
7–10 × 5–7 mm. |
= 16. |
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Flowering Apr–Jul. |
Flowering Apr–Jun. |
Sandy or calcareous soils, tallgrass, mixed-grass, and sand prairies. |
Sagebrush shrublands, pinyon-juniper woodlands. |
200–1800(–2400) m. (700–5900(–7900) ft.) |
400–2500 m. (1300–8200 ft.) |
CO; CT; IA; IL; IN; KS; MA; MI; MN; MO; MT; ND; NE; NM; OH; OK; SD; TX; WY
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AZ; CA; CO; NV; UT
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Widely distributed in the western Midwest and Great Plains in the United States, Penstemon grandiflorus is cultivated as an ornamental for its showy flowers. Reports from Connecticut (D. W. Magee and H. E. Ahles 2007), Indiana (K. Yatskievych 2000), Massachusetts (Magee and Ahles), Michigan (E. G. Voss 1972–1996), and Ohio (T. S. Cooperrider 1995) appear to be based on introductions. Penstemon grandiflorus has been seeded along highways in Iowa and Nebraska, where it is also native. The validity of the name Penstemon grandiflorus has been debated owing to the meager diagnosis by Nuttall. The name is accepted here, making P. bradburyi Pursh, an illegitimate, superfluous replacement for P. grandiflorus. The Dakota, Kiowa, and Sioux tribes, centered in the Great Plains, use Penstemon grandiflorus as an analgesic, a gastrointestinal aid, and for fevers (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon utahensis is widespread in the Colorado River Basin from western Colorado through northwestern Arizona, southeastern California, southern Nevada, and southern Utah. The Hopi tribe of northeastern Arizona used Penstemon utahensis as a ceremonial and decorative plant (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 118. |
FNA vol. 17, p. 155. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
Nuttall: Cat. Pl. Upper Louisiana, no. 64. (1813) — (as grandiflorum) |
Eastwood: Zoë 4: 124. (1893) |
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