large beardtongue, largeflower penstemon, shell-leaf beardtongue
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Rydberg's beardtongue, Rydberg's penstemon
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Herbs. |
erect, (40–)50–95(–120) cm, glabrous. |
decumbent, ascending, or erect, 20–70(–120) cm, glabrous or retrorsely hairy, sometimes only in lines, not glaucous. |
basal and cauline, glabrous; basal and proximal cauline 30–160 × 6–50 mm, blade spatulate to obovate, base tapered, apex rounded to obtuse or acute; cauline 4–8 pairs, sessile, 18–90(–110) × 15–50 mm, blade spatulate to orbiculate, base clasping, apex rounded to obtuse. |
basal and cauline, not leathery, glabrous; basal and proximal cauline (25–)35–70(–150) × (5–)10–15(–22) mm, blade oblanceolate to elliptic, base tapered, margins entire, apex obtuse to acute, rarely mucronate; cauline 3–6 pairs, sessile or proximals short-petiolate, 25–70(–110) × 3–24 mm, blade lanceolate to elliptic or oblong, base tapered to clasping, margins entire, apex obtuse to acute. |
interrupted, cylindric, 12–30(–40) cm, axis glabrous, verticillasters 3–7(–9), cymes 2–4-flowered; proximal bracts ovate to elliptic or orbiculate, (9–)16–83 × (9–)16–54 mm; peduncles and pedicels glabrous. |
interrupted, cylindric, 3–10(–18) cm, axis retrorsely hairy or puberulent, sometimes glabrous, verticillasters (1 or)2–7, cymes (3–)5–11-flowered, 2 per node; proximal bracts lanceolate, (4–)11–52(–70) × (1–)2–18 mm, margins entire or erose to lacerate; peduncles and pedicels erect, glabrous, retrorsely hairy, or puberulent. |
calyx lobes ovate to lanceolate, 7–11 × 2.5–4 mm, margins entire, rarely erose, herbaceous or narrowly scarious, glabrous; corolla lavender to blue or pinkish blue, with magenta nectar guides, ampliate, 35–48 mm, glabrous externally, glabrous internally, tube 10–13 mm, throat abruptly inflated, 15–18 mm diam., rounded abaxially; stamens included, pollen sacs opposite, 2.1–2.6 mm, sutures papillate; staminode 16–21 mm, included or reaching orifice, 2–2.6 mm diam., tip recurved to coiled, distal 1–2 mm sparsely villous, hairs golden yellow, to 0.5 mm; style 19–30 mm. |
calyx lobes oblong to ovate or obovate, 3–8 × 1.2–3.5 mm, apex acute to acuminate or short- to long-caudate, glabrous or puberulent; corolla violet to blue or purple, without nectar guides, funnelform, 10–20 mm, glabrous externally, sparsely white- or yellowish villous internally abaxially, tube 3–4 mm, throat gradually inflated, 3–4.5 mm diam., 2-ridged abaxially; stamens: longer pair reaching orifice or exserted, pollen sacs opposite, navicular, 0.5–1.1 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or ± papillate; staminode 6–7 mm, reaching orifice, 0.2–0.4 mm diam., tip straight, distal 1.5–2 mm moderately to densely pilose, hairs golden yellow, to 1 mm; style 8–10 mm. |
16–20(–25) × 8–15 mm. |
4–5 × 3–4 mm, glabrous. |
= 16. |
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Flowering Apr–Jul. |
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Sandy or calcareous soils, tallgrass, mixed-grass, and sand prairies. |
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200–1800(–2400) m. (700–5900(–7900) ft.) |
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CO; CT; IA; IL; IN; KS; MA; MI; MN; MO; MT; ND; NE; NM; OH; OK; SD; TX; WY
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AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY
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Widely distributed in the western Midwest and Great Plains in the United States, Penstemon grandiflorus is cultivated as an ornamental for its showy flowers. Reports from Connecticut (D. W. Magee and H. E. Ahles 2007), Indiana (K. Yatskievych 2000), Massachusetts (Magee and Ahles), Michigan (E. G. Voss 1972–1996), and Ohio (T. S. Cooperrider 1995) appear to be based on introductions. Penstemon grandiflorus has been seeded along highways in Iowa and Nebraska, where it is also native. The validity of the name Penstemon grandiflorus has been debated owing to the meager diagnosis by Nuttall. The name is accepted here, making P. bradburyi Pursh, an illegitimate, superfluous replacement for P. grandiflorus. The Dakota, Kiowa, and Sioux tribes, centered in the Great Plains, use Penstemon grandiflorus as an analgesic, a gastrointestinal aid, and for fevers (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). As treated here, Penstemon rydbergii includes three partially sympatric varieties. D. D. Keck (1945) treated the same taxa as four species and two subspecies (P. hesperius, P. oreocharis, P. rydbergii subsp. aggregatus, P. rydbergii subsp. rydbergii, and P. vaseyanus). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Calyx lobe margins lacerate, rarely erose, broadly scarious, apices acuminate or short- to long-caudate; proximal bract margins erose to lacerate proximally, sometimes entire; thyrse axes retrorsely hairy or puberulent. | var. rydbergii |
1. Calyx lobe margins erose, sometimes lacerate, narrowly to broadly scarious, apices acute, acuminate, or short-caudate; proximal bract margins entire; thyrse axes glabrous or retrorsely hairy. | → 2 |
2. Stems ± retrorsely hairy; thyrse axes glabrous or retrorsely hairy; corollas (10–)13–20 mm; pollen sacs (0.6–)0.8–1.1 mm. | var. aggregatus |
2. Stems glabrous, rarely ± retrorsely hairy; thyrse axes glabrous, rarely ± retrorsely hairy; corollas 10–14 mm; pollen sacs 0.5–0.7(–0.8) mm. | var. oreocharis |
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FNA vol. 17, p. 118. |
FNA vol. 17, p. 220. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Nuttall: Cat. Pl. Upper Louisiana, no. 64. (1813) — (as grandiflorum) |
A. Nelson: Bull. Torrey Bot. Club 25: 281. (1898) — (as Pentstemon) |
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