large beardtongue, largeflower penstemon, shell-leaf beardtongue
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red desert beardtongue, sand penstemon
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erect, (40–)50–95(–120) cm, glabrous. |
decumbent to ascending, (4–)7–18(–30) cm, glabrous. |
basal and cauline, glabrous; basal and proximal cauline 30–160 × 6–50 mm, blade spatulate to obovate, base tapered, apex rounded to obtuse or acute; cauline 4–8 pairs, sessile, 18–90(–110) × 15–50 mm, blade spatulate to orbiculate, base clasping, apex rounded to obtuse. |
basal and cauline, glabrous; basal and proximal cauline 21–50(–70) × 4–11(–17) mm, blade spatulate to oblanceolate, base tapered, apex rounded to obtuse, acute, or mucronate; cauline 2–5 pairs, short-petiolate or sessile, 21–58 × 3–11 mm, blade lanceolate to oblanceolate, base tapered to clasping, apex obtuse to acute or mucronate. |
interrupted, cylindric, 12–30(–40) cm, axis glabrous, verticillasters 3–7(–9), cymes 2–4-flowered; proximal bracts ovate to elliptic or orbiculate, (9–)16–83 × (9–)16–54 mm; peduncles and pedicels glabrous. |
continuous, cylindric, 3–14 cm, axis glabrous, verticillasters 4–9, cymes (1 or)2–6-flowered; proximal bracts oblanceolate to lanceolate, 13–60 × 5–16 mm; peduncles and pedicels glabrous. |
calyx lobes ovate to lanceolate, 7–11 × 2.5–4 mm, margins entire, rarely erose, herbaceous or narrowly scarious, glabrous; corolla lavender to blue or pinkish blue, with magenta nectar guides, ampliate, 35–48 mm, glabrous externally, glabrous internally, tube 10–13 mm, throat abruptly inflated, 15–18 mm diam., rounded abaxially; stamens included, pollen sacs opposite, 2.1–2.6 mm, sutures papillate; staminode 16–21 mm, included or reaching orifice, 2–2.6 mm diam., tip recurved to coiled, distal 1–2 mm sparsely villous, hairs golden yellow, to 0.5 mm; style 19–30 mm. |
calyx lobes lanceolate, 4–6.2(–7.5) × 0.9–1.6 mm, margins entire or erose, narrowly scarious, glabrous; corolla blue to violet, with reddish purple nectar guides, tubular-funnelform, (10–)12–15 mm, glabrous externally, sparsely white-pubescent internally abaxially or glabrous, tube 4–5 mm, throat gradually inflated, 4.5–5.5 mm diam., rounded abaxially; stamens included, pollen sacs divergent, 0.7–0.8 mm, sutures smooth or obscurely papillate; staminode 6–8 mm, included, 0.8–1 mm diam., tip recurved, distal 3–4 mm densely pubescent, hairs golden yellow, to 1 mm; style 8–10 mm. |
16–20(–25) × 8–15 mm. |
6–10(–12) × 5–6 mm. |
= 16. |
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Flowering Apr–Jul. |
Flowering May–Jun. |
Sandy or calcareous soils, tallgrass, mixed-grass, and sand prairies. |
Sandy or shaley soils, sagebrush shrublands. |
200–1800(–2400) m. (700–5900(–7900) ft.) |
1800–2400 m. (5900–7900 ft.) |
CO; CT; IA; IL; IN; KS; MA; MI; MN; MO; MT; ND; NE; NM; OH; OK; SD; TX; WY
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WY
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Widely distributed in the western Midwest and Great Plains in the United States, Penstemon grandiflorus is cultivated as an ornamental for its showy flowers. Reports from Connecticut (D. W. Magee and H. E. Ahles 2007), Indiana (K. Yatskievych 2000), Massachusetts (Magee and Ahles), Michigan (E. G. Voss 1972–1996), and Ohio (T. S. Cooperrider 1995) appear to be based on introductions. Penstemon grandiflorus has been seeded along highways in Iowa and Nebraska, where it is also native. The validity of the name Penstemon grandiflorus has been debated owing to the meager diagnosis by Nuttall. The name is accepted here, making P. bradburyi Pursh, an illegitimate, superfluous replacement for P. grandiflorus. The Dakota, Kiowa, and Sioux tribes, centered in the Great Plains, use Penstemon grandiflorus as an analgesic, a gastrointestinal aid, and for fevers (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon arenicola occurs throughout much of the western two-thirds of Wyoming. Specimens documenting occurrence of the species in Colorado and Montana have not been seen. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 118. |
FNA vol. 17, p. 116. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
Nuttall: Cat. Pl. Upper Louisiana, no. 64. (1813) — (as grandiflorum) |
A. Nelson: Bull. Torrey Bot. Club 25: 280. (1898) — (as Pentstemon) |
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