glandular penstemon, sticky-stem beardtongue, sticky-stem penstemon
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Penland's beardtongue
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Herbs. |
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ascending to erect, (30–)40–100 cm, viscid glandular-pubescent, not glaucous. |
ascending to erect, 7–25 cm, retrorsely hairy or puberulent, not glaucous. |
basal and cauline, opposite, viscid glandular-pubescent, not glaucous; basal and proximal cauline 25–120(–260) × 7–20(–65) mm, blade elliptic to lanceolate, base tapered, margins entire, subentire, or irregularly serrate to dentate, apex acute to acuminate; cauline 4–10 pairs, sessile, 32–80(–120) × 12–45 mm, blade ovate to obovate, elliptic, or lanceolate, base tapered to cordate-clasping, margins entire or subentire to irregularly serrate to dentate, apex obtuse to acute or acuminate. |
basal and cauline, not leathery, puberulent or retrorsely hairy, not glaucous; basal and proximal cauline 15–80 × 0.8–2 mm, blade linear, base tapered, margins entire, apex obtuse to acute; cauline 1–3 pairs, sessile, 10–75 × 0.5–1.5 mm, blade linear, base tapered, apex acute. |
interrupted, cylindric, (5–)8–31 cm, axis viscid glandular-pubescent, verticillasters 3–7, cymes (1 or)2–8-flowered, 2 per node; proximal bracts ovate, 18–60 × 11–44 mm; peduncles and pedicels ascending to erect, viscid glandular-pubescent. |
continuous, secund, 3–10 cm, axis glandular-pubescent, verticillasters 2–6, cymes 1- or 2(–4)-flowered, 2 per node; proximal bracts linear, 10–56 × 0.5–1.5 mm; peduncles and pedicels glandular-pubescent. |
calyx lobes lanceolate, 9–15 × 2.3–3.8 mm, viscid glandular-pubescent; corolla lavender to bluish lavender or purple, with dark violet nectar guides, ampliate, (28–)30–40 mm, glandular-pubescent externally, sparsely to moderately white-pubescent internally abaxially, tube 9–11 mm, throat gradually inflated, 12–15 mm diam., rounded abaxially; stamens included, filaments glabrous, pollen sacs parallel, 1.6–2.2 mm, distal 1/2 indehiscent, sides glabrous, sutures denticulate, teeth to 0.2 mm; staminode 22–26 mm, exserted, 0.8–1 mm diam., glabrous, rarely distal 2–4 mm sparsely pubescent, hairs yellow, to 0.4 mm; style 20–26 mm. |
calyx lobes lanceolate, 4.5–5.5 × 1.8–2.5 mm, glabrous to sparsely glandular; corolla blue to bluish violet, with reddish purple nectar guides, funnelform, 13–16 mm, glabrous externally, glabrous internally, tube 5–6 mm, throat gradually inflated, not constricted at orifice, 4.5–5.5 mm diam., rounded abaxially; stamens included, pollen sacs divergent to opposite, navicular, 1.1–1.5 mm, dehiscing incompletely, connective not splitting, sides moderately hirsute, hairs white, to 0.5 mm, sutures denticulate, teeth to 0.1 mm; staminode 8–9 mm, included, 0.9–1.1 mm diam., tip straight, distal 6–7 mm densely pubescent, hairs yellowish orange, to 0.9 mm; style 8–10 mm. |
9–13 × 5–6.5 mm. |
6–9(–14) × 4–6 mm. |
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Flowering Jun–Jul. |
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Clayey soils, sagebrush shrublands. |
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2300–2400 m. (7500–7900 ft.) |
ID; OR; WA
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CO |
Varieties 2 (2 in the flora). Two allopatric varieties of Penstemon glandulosus have been distinguished based on basal leaf morphology. Leaf margins occasionally vary from entire to serrate or dentate among individuals in some populations, but the general pattern of leaf variation does appear to be geographically correlated. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon penlandii is known from Middle Park in Grand County. Plants grow on strongly seleniferous shales of the Troublesome Formation (S. L. O’Kane 1988). The species is listed as endangered by the U.S. Department of the Interior. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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FNA vol. 17, p. 233. |
FNA vol. 17, p. 176. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1262. (1829) — (as Pentstemon glandulosum) |
W. A. Weber: Phytologia 60: 459. (1986) |
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