glandular penstemon, sticky-stem beardtongue, sticky-stem penstemon
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Guadalupe beardtongue
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Herbs. |
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ascending to erect, (30–)40–100 cm, viscid glandular-pubescent, not glaucous. |
ascending to erect, 20–50 cm, retrorsely hairy. |
basal and cauline, opposite, viscid glandular-pubescent, not glaucous; basal and proximal cauline 25–120(–260) × 7–20(–65) mm, blade elliptic to lanceolate, base tapered, margins entire, subentire, or irregularly serrate to dentate, apex acute to acuminate; cauline 4–10 pairs, sessile, 32–80(–120) × 12–45 mm, blade ovate to obovate, elliptic, or lanceolate, base tapered to cordate-clasping, margins entire or subentire to irregularly serrate to dentate, apex obtuse to acute or acuminate. |
basal and cauline, not leathery, glabrous or glabrate, rarely basal retrorsely hairy along midvein; basal and proximal cauline petiolate, 25–95 × 1–6 mm, blade linear, rarely lanceolate, base tapered, margins entire, apex acute; cauline 6–11, sessile or proximals short-petiolate, 22–65(–95) × 1–10(–18) mm, blade lanceolate to linear, base tapered to clasping, margins entire or obscurely serrate, apex acute. |
interrupted, cylindric, (5–)8–31 cm, axis viscid glandular-pubescent, verticillasters 3–7, cymes (1 or)2–8-flowered, 2 per node; proximal bracts ovate, 18–60 × 11–44 mm; peduncles and pedicels ascending to erect, viscid glandular-pubescent. |
continuous or ± interrupted, cylindric, 3–16(–26) cm, axis glandular-pubescent, verticillasters 3–8, cymes 2–4-flowered, 2 per node; proximal bracts lanceolate, 15–48 × 3–8 mm; peduncles and pedicels glandular-pubescent. |
calyx lobes lanceolate, 9–15 × 2.3–3.8 mm, viscid glandular-pubescent; corolla lavender to bluish lavender or purple, with dark violet nectar guides, ampliate, (28–)30–40 mm, glandular-pubescent externally, sparsely to moderately white-pubescent internally abaxially, tube 9–11 mm, throat gradually inflated, 12–15 mm diam., rounded abaxially; stamens included, filaments glabrous, pollen sacs parallel, 1.6–2.2 mm, distal 1/2 indehiscent, sides glabrous, sutures denticulate, teeth to 0.2 mm; staminode 22–26 mm, exserted, 0.8–1 mm diam., glabrous, rarely distal 2–4 mm sparsely pubescent, hairs yellow, to 0.4 mm; style 20–26 mm. |
calyx lobes lanceolate, 5–6.5 × 1.2–2 mm, glandular-pubescent; corolla white, sometimes tinged pink or lavender, with reddish violet nectar guides, funnelform, 13–20 mm, glandular-pubescent and sparsely white-villous internally abaxially, tube 4–6 mm, throat gradually inflated, not constricted at orifice, 6–7 mm diam., rounded abaxially; stamens included, pollen sacs opposite, explanate, 0.8–1 mm, dehiscing completely, sutures smooth; staminode 10–14 mm, reaching orifice or exserted, 0.5–0.7 mm diam., tip straight to recurved, distal 3–7 mm sparsely pilose, hairs yellow, to 0.9 mm; style 9–12 mm. |
9–13 × 5–6.5 mm. |
7–9 × 4.5–6 mm. |
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Flowering Mar–Jun. |
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Sandy or gravelly, calcareous soils, prairies. |
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400–800 m. (1300–2600 ft.) |
ID; OR; WA
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TX |
Varieties 2 (2 in the flora). Two allopatric varieties of Penstemon glandulosus have been distinguished based on basal leaf morphology. Leaf margins occasionally vary from entire to serrate or dentate among individuals in some populations, but the general pattern of leaf variation does appear to be geographically correlated. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon guadalupensis is known from the southern Cross Timbers and Prairies, Edwards Plateau, and southern Rolling Plains physiographic provinces of central Texas. Pennell distinguished subsp. ernestii by its shorter corollas and broader leaves, but that distinction seems trivial. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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FNA vol. 17, p. 233. |
FNA vol. 17, p. 137. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. guadalupensis subsp. ernestii |
Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1262. (1829) — (as Pentstemon glandulosum) |
A. Heller: Contr. Herb. Franklin Marshall Coll. 1: 92, plate 7. (1895) — (as Pentstemon) |
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