glandular penstemon, sticky-stem beardtongue, sticky-stem penstemon
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short-sepal beardtongue
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Herbs. |
Herbs. |
ascending to erect, (30–)40–100 cm, viscid glandular-pubescent, not glaucous. |
ascending to erect, 22–60 cm, retrorsely hairy and glandular-pubescent or glandular-villous, not glaucous. |
basal and cauline, opposite, viscid glandular-pubescent, not glaucous; basal and proximal cauline 25–120(–260) × 7–20(–65) mm, blade elliptic to lanceolate, base tapered, margins entire, subentire, or irregularly serrate to dentate, apex acute to acuminate; cauline 4–10 pairs, sessile, 32–80(–120) × 12–45 mm, blade ovate to obovate, elliptic, or lanceolate, base tapered to cordate-clasping, margins entire or subentire to irregularly serrate to dentate, apex obtuse to acute or acuminate. |
basal and cauline, basal sometimes withering by anthesis, not leathery, sparsely to densely pubescent, usually also sparsely glandular-hairy, abaxially, glabrous or sparsely pubescent adaxially; basal and proximal cauline 20–95 × 11–28 mm, blade spatulate to obovate or ovate, base tapered, margins subentire to ± serrate, apex rounded to obtuse; cauline 4 or 5, sessile or proximals short-petiolate, 35–95 × 11–32 mm, blade ovate to lanceolate or oblanceolate, base tapered to truncate or clasping, margins ± serrate or dentate, apex obtuse to acute. |
interrupted, cylindric, (5–)8–31 cm, axis viscid glandular-pubescent, verticillasters 3–7, cymes (1 or)2–8-flowered, 2 per node; proximal bracts ovate, 18–60 × 11–44 mm; peduncles and pedicels ascending to erect, viscid glandular-pubescent. |
interrupted, conic, (4–)8–20 cm, axis sparsely to densely glandular-pubescent, verticillasters 3–8, cymes 5–16-flowered, 2 per node; proximal bracts lanceolate, 7–50 × 2–20 mm, margins entire or serrate; peduncles and pedicles spreading to ascending, sparsely to densely glandular-pubescent. |
calyx lobes lanceolate, 9–15 × 2.3–3.8 mm, viscid glandular-pubescent; corolla lavender to bluish lavender or purple, with dark violet nectar guides, ampliate, (28–)30–40 mm, glandular-pubescent externally, sparsely to moderately white-pubescent internally abaxially, tube 9–11 mm, throat gradually inflated, 12–15 mm diam., rounded abaxially; stamens included, filaments glabrous, pollen sacs parallel, 1.6–2.2 mm, distal 1/2 indehiscent, sides glabrous, sutures denticulate, teeth to 0.2 mm; staminode 22–26 mm, exserted, 0.8–1 mm diam., glabrous, rarely distal 2–4 mm sparsely pubescent, hairs yellow, to 0.4 mm; style 20–26 mm. |
calyx lobes ovate, rarely lanceolate, (1.5–)2–3.5 × 0.9–2.2 mm, sparsely glandular-pubescent; corolla lavender to purple, with reddish purple nectar guides, tubular to tubular-funnelform, 15–24 mm, glandular-pubescent externally, ± whitish lanate internally abaxially, tube 4–6 mm, throat slightly inflated, 4–7 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular, 0.9–1.1 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 12–18 mm, exserted, 0.3–0.5 mm diam., tip straight to slightly recurved, distal 8–9 mm ± villous, hairs yellow or golden yellow, to 1.5 mm; style 12–17 mm. |
9–13 × 5–6.5 mm. |
4–7 × 3–5 mm, glabrous. |
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Flowering May–Jun. |
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Rocky oak-hickory forests, bluffs, roadcuts. |
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200–700 m. (700–2300 ft.) |
ID; OR; WA
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KY; TN; VA; WV |
Varieties 2 (2 in the flora). Two allopatric varieties of Penstemon glandulosus have been distinguished based on basal leaf morphology. Leaf margins occasionally vary from entire to serrate or dentate among individuals in some populations, but the general pattern of leaf variation does appear to be geographically correlated. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon brevisepalus has not been widely recognized in the literature or herbaria, usually being synonymized with P. canescens or P. pallidus. Penstemon brevisepalus combines features of P. canescens and P. pallidus, and its range lies between the ranges of those two species, with little overlap. In addition to calyx lobe length and corolla color, P. brevisepalus usually has more open inflorescences and cauline leaf margins that are more finely and regularly toothed compared to P. pallidus. Plants with pink- or lavender-tinged corollas occasionally occur in P. pallidus, and individuals with short calyx lobes (2–3 mm) also are encountered infrequently; they may be the sources of reports of P. brevisepalus elsewhere (for example, Georgia, Illinois, and Ohio). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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FNA vol. 17, p. 233. |
FNA vol. 17, p. 196. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1262. (1829) — (as Pentstemon glandulosum) |
Pennell: in J. K. Small, Man. S.E. Fl., 1204, 1508. (1933) |
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