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glandular penstemon, sticky-stem beardtongue, sticky-stem penstemon

Nevada sand-dune beardtongue

Habit Herbs. Herbs.
Caudex

rhizomelike.

Stems

ascending to erect, (30–)40–100 cm, viscid glandular-pubescent, not glaucous.

ascending to erect, 10–27 cm, glabrous, glutinous, usually covered with sand, not glaucous.

Leaves

basal and cauline, opposite, viscid glandular-pubescent, not glaucous;

basal and proximal cauline 25–120(–260) × 7–20(–65) mm, blade elliptic to lanceolate, base tapered, margins entire, subentire, or irregularly serrate to dentate, apex acute to acuminate;

cauline 4–10 pairs, sessile, 32–80(–120) × 12–45 mm, blade ovate to obovate, elliptic, or lanceolate, base tapered to cordate-clasping, margins entire or subentire to irregularly serrate to dentate, apex obtuse to acute or acuminate.

cauline, leathery, glabrous, sometimes glutinous;

cauline 3–5 pairs, short-petiolate or sessile, (12–)20–50(–55) × (3–)10–24 mm, blade obovate or spatulate proximally, obovate to lanceolate distally, base tapered, margins dentate, apex rounded to obtuse or acute.

Thyrses

interrupted, cylindric, (5–)8–31 cm, axis viscid glandular-pubescent, verticillasters 3–7, cymes (1 or)2–8-flowered, 2 per node;

proximal bracts ovate, 18–60 × 11–44 mm;

peduncles and pedicels ascending to erect, viscid glandular-pubescent.

continuous, cylindric, (1–)2–9 cm, axis glabrous or sparsely puberulent, glutinous, verticillasters 2–8, cymes 2–6-flowered, 2 per node;

proximal bracts lanceolate, 16–58 × 3–12 mm, margins dentate;

peduncles and pedicels erect, glabrous or puberulent, glutinous.

Flowers

calyx lobes lanceolate, 9–15 × 2.3–3.8 mm, viscid glandular-pubescent;

corolla lavender to bluish lavender or purple, with dark violet nectar guides, ampliate, (28–)30–40 mm, glandular-pubescent externally, sparsely to moderately white-pubescent internally abaxially, tube 9–11 mm, throat gradually inflated, 12–15 mm diam., rounded abaxially;

stamens included, filaments glabrous, pollen sacs parallel, 1.6–2.2 mm, distal 1/2 indehiscent, sides glabrous, sutures denticulate, teeth to 0.2 mm;

staminode 22–26 mm, exserted, 0.8–1 mm diam., glabrous, rarely distal 2–4 mm sparsely pubescent, hairs yellow, to 0.4 mm;

style 20–26 mm.

calyx lobes ovate to lanceolate, 4.5–8.4 × 1.6–3 mm, glutinous;

corolla white to lavender, with reddish purple nectar guides, tubular-funnelform, 11–15(–18) mm, glabrous externally, sparsely white-pilose internally abaxially, tube 6–7 mm, throat gradually inflated, 3–5 mm diam., rounded abaxially;

stamens included, pollen sacs opposite, navicular to subexplanate, 0.8–0.9 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth;

staminode 7–10 mm, included, 0.6–0.8 mm diam., tip slightly recurved, distal 0.5–2.5 mm sparsely pilose, hairs white, to 0.5 mm;

style 7–8 mm.

Capsules

9–13 × 5–6.5 mm.

5–8 × 4–5 mm, glabrous.

2n

= 16.

Penstemon glandulosus

Penstemon arenarius

Phenology Flowering May–Jul.
Habitat Loose sands, dunes, valleys, desert shrublands.
Elevation 1200–1800 m. (3900–5900 ft.)
Distribution
from FNA
ID; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
NV
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Two allopatric varieties of Penstemon glandulosus have been distinguished based on basal leaf morphology. Leaf margins occasionally vary from entire to serrate or dentate among individuals in some populations, but the general pattern of leaf variation does appear to be geographically correlated.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon arenarius has been documented in Churchill, Mineral, and Nye counties. Plants usually are covered with sand because of their glutinous herbage.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves: blade margins irregularly serrate to dentate.
var. glandulosus
1. Basal leaves: blade margins entire or subentire.
var. chelanensis
Source FNA vol. 17, p. 233. FNA vol. 17, p. 191.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. glandulosus var. chelanensis, P. glandulosus var. glandulosus
Name authority Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1262. (1829) — (as Pentstemon glandulosum) Greene: Pittonia 1: 282. (1889) — (as Pentstemon)
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