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sawsepal penstemon, southern smooth beardtongue, western smooth beardtongue

Apache beardtongue

Habit Herbs.
Stems

ascending, (10–)50–65(–80) cm, glabrous or puberulent to pubescent, not glaucous.

erect, (6–)10–45 cm, retrorsely hairy, also glandular-pubescent distally, not glaucous.

Leaves

basal and cauline, or basal absent or reduced, not leathery, glabrous or puberulent to pubescent, not glaucous;

basal and proximal cauline 20–80(–155) × 5–20(–45) mm, blade obovate to oblanceolate or lanceolate, base tapered, margins entire, usually undulate, apex obtuse to acute, sometimes mucronate;

cauline (2 or)3–6(–8) pairs, sessile, 27–120(–150) × (6–)22–35(–43) mm, blade lanceolate, base truncate to cordate, apex obtuse to acute.

basal and cauline, not leathery, proximals usually retrorsely hairy proximally, especially along midveins and margins, mostly glabrate distally, distals retrorsely hairy and, sometimes, sparsely glandular-pubescent, sometimes ± glaucescent;

basal and proximal cauline 15–75 × (2–)5–19 mm, blade spatulate to oblanceolate or elliptic, base tapered, margins entire, apex rounded to obtuse or acute, sometimes mucronate;

cauline 2–5 pairs, sessile, 11–70 × (1–)4–15 mm, blade of proximal leaves oblanceolate to lanceolate, distal leaves lanceolate, base tapered to clasping, margins entire, rarely ± denticulate distally, apex acute to acuminate.

Thyrses

continuous, secund, (3–)6–26(–30) cm, axis glabrous or puberulent, verticillasters (3–)5–12, cymes 2–4-flowered, 2 per node;

proximal bracts ovate to lanceolate, 14–100 × 2–36 mm;

peduncles and pedicels glabrous or puberulent, peduncles to 71 mm, pedicels 4–14 mm.

interrupted, secund, (1–)3–24(–30) cm, axis glandular-pubescent, verticillasters (2 or)3–7, cymes 1–4-flowered, (1 or)2 per node;

proximal bracts lanceolate to linear, 7–38(–60) × 1–7 mm, margins entire;

peduncles and pedicels erect, glandular-pubescent.

Flowers

calyx lobes ovate to orbiculate, 2–11 × 1.3–5.5 mm, glabrous or puberulent;

corolla blue to violet, with reddish purple nectar guides, ventricose, 24–48 mm, glabrous externally, glabrous internally or sparsely to moderately white-lanate internally abaxially, tube 5–12 mm, throat gradually inflated, slightly constricted at orifice, 8–13(–18) mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, pollen sacs divergent, navicular, 1.4–2.5(–2.8) mm, dehiscing incompletely, connective not splitting, sides sparsely to moderately hirsute, hairs white or tan, to 0.5 mm, rarely glabrous, sutures denticulate, teeth to 0.2 mm;

staminode (15–)18–22 mm, included or barely exserted, 0.6–1.3 mm diam., tip straight to recurved, rounded to distinctly bifurcate at apex, glabrous or distal 1–2 mm sparsely lanate, hairs pale yellow, to 1.5 mm;

style 18–20 mm.

calyx lobes lanceolate, 2.8–6 × 0.9–2 mm, glandular-pubescent;

corolla blue to violet-blue, with reddish purple nectar guides, tubular to tubular-funnelform, 11–20 mm, glandular-pubescent externally, sparsely to moderately white-villous to white-lanate internally abaxially, tube 5–7 mm, throat slightly inflated, 3–6 mm diam., 2-ridged abaxially;

stamens included, pollen sacs opposite, navicular, 0.8–1.2 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate;

staminode 10–12 mm, included or slightly exserted, 0.3–0.4 mm diam., tip straight to slightly recurved, distal 3–6 mm densely pilose, hairs golden, to 2 mm;

style 9–13 mm.

Capsules

10–17 × 5–8 mm.

5–7 × 3–4.5 mm, glabrous.

2n

= 16.

Penstemon glaber

Penstemon oliganthus

Phenology Flowering Jun–Aug(–Oct).
Habitat Montane meadows, ciénagas, clearings in pine and spruce-fir forests.
Elevation 2400–3500 m. (7900–11500 ft.)
Distribution
from FNA
CO; MT; ND; NE; NM; SD; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

The Penstemon glaber complex has been treated with as many as four species (F. W. Pennell 1920b) or one variable species as treated herein. M. K. W. Latady (1985) found clinal variation in most morphologic characters used to distinguish taxa in the complex and recognized only one species and two varieties: var. glaber (including var. alpinus) in the plains and mountains from southeastern Montana south to central Colorado; and var. brandegeei, in the mountains from central Colorado south to north-central New Mexico. A case for recognizing var. alpinus, as is done here, can be made on morphological, phenological, and ecological grounds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon oliganthus is known from the Mogollon Rim (Coconino County) and White Mountains of east-central Arizona (Apache and Greenlee counties) and the Jemez and San Mateo mountains of northwestern New Mexico (McKinley, Sandoval, and Valencia counties). Crosswhite described P. pseudoparvus from five specimens from the Magdalena and San Mateo mountains in Socorro County, New Mexico, and more than 24 specimens of P. oliganthus from Arizona and New Mexico. He separated the two species based on stem indument (obscurely puberulent in P. oliganthus versus obviously puberulent in P. pseudoparvus), and flower orientation and shape (horizontal or more usually drooping and little inflated in P. oliganthus versus ascending and not inflated in P. pseudoparvus). More than 60 collections referable to these species were examined for this treatment, including the types, specimens annotated by Crosswhite, and 20 specimens from the San Mateo and Magdalena mountains in Socorro County; the two taxa appear to be indistinguishable.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx lobes 2–4.8 mm, apices rounded or abruptly short-acuminate; staminode apices rounded.
var. glaber
1. Calyx lobes 3.8–11 mm, apices long-acuminate; staminode apices rounded or bifurcate.
→ 2
2. Staminode apices rounded or bifurcate; corollas 24–35 mm; stems glabrous or puberulent proximally.
var. alpinus
2. Staminode apices usually bifurcate; corollas 27–48 mm; stems puberulent to pubescent proximally.
var. brandegeei
Source FNA vol. 17, p. 167. FNA vol. 17, p. 213.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. glaber var. alpinus, P. glaber var. brandegeei, P. glaber var. glaber
Synonyms P. pseudoparvus
Name authority Pursh: Fl. Amer. Sept. 2: 738. (1813) — (as Pentstemon glabra) Wooton & Standley: Contr. U.S. Natl. Herb. 16: 172. (1913) — (as Pentstemon)
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