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sawsepal penstemon, southern smooth beardtongue, western smooth beardtongue

golden tongue penstemon, Malheur beardtongue, Malheur penstemon

Stems

ascending, (10–)50–65(–80) cm, glabrous or puberulent to pubescent, not glaucous.

ascending to erect, 8–28(–40) cm, retrorsely hairy.

Leaves

basal and cauline, or basal absent or reduced, not leathery, glabrous or puberulent to pubescent, not glaucous;

basal and proximal cauline 20–80(–155) × 5–20(–45) mm, blade obovate to oblanceolate or lanceolate, base tapered, margins entire, usually undulate, apex obtuse to acute, sometimes mucronate;

cauline (2 or)3–6(–8) pairs, sessile, 27–120(–150) × (6–)22–35(–43) mm, blade lanceolate, base truncate to cordate, apex obtuse to acute.

basal and cauline, not leathery, retrorsely hairy, hairs sometimes appressed, white, scalelike, sometimes basal and proximal cauline glabrate or glabrous abaxially;

basal and proximal cauline petiolate, 15–60(–78) × 4–12(–17) mm, blade spatulate to oblanceolate, base tapered, margins entire or dentate-serrate, apex rounded to obtuse or acute;

cauline 3 or 4 pairs, sessile or proximals short-petiolate, 20–50(–75) × 2–5(–10) mm, blade lanceolate to oblanceolate, base tapered to clasping, margins entire or obscurely and remotely dentate, apex obtuse to acute.

Thyrses

continuous, secund, (3–)6–26(–30) cm, axis glabrous or puberulent, verticillasters (3–)5–12, cymes 2–4-flowered, 2 per node;

proximal bracts ovate to lanceolate, 14–100 × 2–36 mm;

peduncles and pedicels glabrous or puberulent, peduncles to 71 mm, pedicels 4–14 mm.

± interrupted, cylindric, (3–)6–18 cm, axis glandular-pubescent and, sometimes, retrorsely hairy, verticillasters 3–6, cymes 2–5-flowered, 2 per node;

proximal bracts lanceolate, 18–46 × 2–12 mm;

peduncles and pedicels glandular-pubescent.

Flowers

calyx lobes ovate to orbiculate, 2–11 × 1.3–5.5 mm, glabrous or puberulent;

corolla blue to violet, with reddish purple nectar guides, ventricose, 24–48 mm, glabrous externally, glabrous internally or sparsely to moderately white-lanate internally abaxially, tube 5–12 mm, throat gradually inflated, slightly constricted at orifice, 8–13(–18) mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, pollen sacs divergent, navicular, 1.4–2.5(–2.8) mm, dehiscing incompletely, connective not splitting, sides sparsely to moderately hirsute, hairs white or tan, to 0.5 mm, rarely glabrous, sutures denticulate, teeth to 0.2 mm;

staminode (15–)18–22 mm, included or barely exserted, 0.6–1.3 mm diam., tip straight to recurved, rounded to distinctly bifurcate at apex, glabrous or distal 1–2 mm sparsely lanate, hairs pale yellow, to 1.5 mm;

style 18–20 mm.

calyx lobes lanceolate, 5.5–8 × 1.2–2 mm, glandular-pubescent;

corolla light blue to bluish purple or magenta, with reddish violet nectar guides, ventricose, 14–22 mm, sparsely white-lanate internally abaxially, tube 5–7 mm, throat abruptly inflated, not constricted at orifice, (3–)5–7 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, pollen sacs opposite, explanate, 0.7–1(–1.2) mm, dehiscing completely, sutures smooth;

staminode 8–9 mm, exserted, 0.3–0.5 mm diam., tip straight to recurved, distal 5–7 mm moderately to densely pilose, hairs yellow or orangish yellow, to 1.2 mm;

style 9–10 mm.

Capsules

10–17 × 5–8 mm.

(6–)8–11 × 4.5–7 mm.

Penstemon glaber

Penstemon miser

Phenology Flowering May–Jun.
Habitat Clayey soils, sagebrush shrublands, pine-juniper woodlands.
Elevation 800–1400 m. (2600–4600 ft.)
Distribution
from FNA
CO; MT; ND; NE; NM; SD; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
ID; OR
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

The Penstemon glaber complex has been treated with as many as four species (F. W. Pennell 1920b) or one variable species as treated herein. M. K. W. Latady (1985) found clinal variation in most morphologic characters used to distinguish taxa in the complex and recognized only one species and two varieties: var. glaber (including var. alpinus) in the plains and mountains from southeastern Montana south to central Colorado; and var. brandegeei, in the mountains from central Colorado south to north-central New Mexico. A case for recognizing var. alpinus, as is done here, can be made on morphological, phenological, and ecological grounds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon miser is known from Owyhee County, Idaho, and southern Baker and northern Malheur counties, Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx lobes 2–4.8 mm, apices rounded or abruptly short-acuminate; staminode apices rounded.
var. glaber
1. Calyx lobes 3.8–11 mm, apices long-acuminate; staminode apices rounded or bifurcate.
→ 2
2. Staminode apices rounded or bifurcate; corollas 24–35 mm; stems glabrous or puberulent proximally.
var. alpinus
2. Staminode apices usually bifurcate; corollas 27–48 mm; stems puberulent to pubescent proximally.
var. brandegeei
Source FNA vol. 17, p. 167. FNA vol. 17, p. 140.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. glaber var. alpinus, P. glaber var. brandegeei, P. glaber var. glaber
Name authority Pursh: Fl. Amer. Sept. 2: 738. (1813) — (as Pentstemon glabra) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. ed. 2, 2(1): 441. (1886) — (as Pentstemon)
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