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sawsepal penstemon, southern smooth beardtongue, western smooth beardtongue

Guadalupe beardtongue

Stems

ascending, (10–)50–65(–80) cm, glabrous or puberulent to pubescent, not glaucous.

ascending to erect, 20–50 cm, retrorsely hairy.

Leaves

basal and cauline, or basal absent or reduced, not leathery, glabrous or puberulent to pubescent, not glaucous;

basal and proximal cauline 20–80(–155) × 5–20(–45) mm, blade obovate to oblanceolate or lanceolate, base tapered, margins entire, usually undulate, apex obtuse to acute, sometimes mucronate;

cauline (2 or)3–6(–8) pairs, sessile, 27–120(–150) × (6–)22–35(–43) mm, blade lanceolate, base truncate to cordate, apex obtuse to acute.

basal and cauline, not leathery, glabrous or glabrate, rarely basal retrorsely hairy along midvein;

basal and proximal cauline petiolate, 25–95 × 1–6 mm, blade linear, rarely lanceolate, base tapered, margins entire, apex acute;

cauline 6–11, sessile or proximals short-petiolate, 22–65(–95) × 1–10(–18) mm, blade lanceolate to linear, base tapered to clasping, margins entire or obscurely serrate, apex acute.

Thyrses

continuous, secund, (3–)6–26(–30) cm, axis glabrous or puberulent, verticillasters (3–)5–12, cymes 2–4-flowered, 2 per node;

proximal bracts ovate to lanceolate, 14–100 × 2–36 mm;

peduncles and pedicels glabrous or puberulent, peduncles to 71 mm, pedicels 4–14 mm.

continuous or ± interrupted, cylindric, 3–16(–26) cm, axis glandular-pubescent, verticillasters 3–8, cymes 2–4-flowered, 2 per node;

proximal bracts lanceolate, 15–48 × 3–8 mm;

peduncles and pedicels glandular-pubescent.

Flowers

calyx lobes ovate to orbiculate, 2–11 × 1.3–5.5 mm, glabrous or puberulent;

corolla blue to violet, with reddish purple nectar guides, ventricose, 24–48 mm, glabrous externally, glabrous internally or sparsely to moderately white-lanate internally abaxially, tube 5–12 mm, throat gradually inflated, slightly constricted at orifice, 8–13(–18) mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, pollen sacs divergent, navicular, 1.4–2.5(–2.8) mm, dehiscing incompletely, connective not splitting, sides sparsely to moderately hirsute, hairs white or tan, to 0.5 mm, rarely glabrous, sutures denticulate, teeth to 0.2 mm;

staminode (15–)18–22 mm, included or barely exserted, 0.6–1.3 mm diam., tip straight to recurved, rounded to distinctly bifurcate at apex, glabrous or distal 1–2 mm sparsely lanate, hairs pale yellow, to 1.5 mm;

style 18–20 mm.

calyx lobes lanceolate, 5–6.5 × 1.2–2 mm, glandular-pubescent;

corolla white, sometimes tinged pink or lavender, with reddish violet nectar guides, funnelform, 13–20 mm, glandular-pubescent and sparsely white-villous internally abaxially, tube 4–6 mm, throat gradually inflated, not constricted at orifice, 6–7 mm diam., rounded abaxially;

stamens included, pollen sacs opposite, explanate, 0.8–1 mm, dehiscing completely, sutures smooth;

staminode 10–14 mm, reaching orifice or exserted, 0.5–0.7 mm diam., tip straight to recurved, distal 3–7 mm sparsely pilose, hairs yellow, to 0.9 mm;

style 9–12 mm.

Capsules

10–17 × 5–8 mm.

7–9 × 4.5–6 mm.

Penstemon glaber

Penstemon guadalupensis

Phenology Flowering Mar–Jun.
Habitat Sandy or gravelly, calcareous soils, prairies.
Elevation 400–800 m. (1300–2600 ft.)
Distribution
from FNA
CO; MT; ND; NE; NM; SD; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

The Penstemon glaber complex has been treated with as many as four species (F. W. Pennell 1920b) or one variable species as treated herein. M. K. W. Latady (1985) found clinal variation in most morphologic characters used to distinguish taxa in the complex and recognized only one species and two varieties: var. glaber (including var. alpinus) in the plains and mountains from southeastern Montana south to central Colorado; and var. brandegeei, in the mountains from central Colorado south to north-central New Mexico. A case for recognizing var. alpinus, as is done here, can be made on morphological, phenological, and ecological grounds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon guadalupensis is known from the southern Cross Timbers and Prairies, Edwards Plateau, and southern Rolling Plains physiographic provinces of central Texas. Pennell distinguished subsp. ernestii by its shorter corollas and broader leaves, but that distinction seems trivial.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx lobes 2–4.8 mm, apices rounded or abruptly short-acuminate; staminode apices rounded.
var. glaber
1. Calyx lobes 3.8–11 mm, apices long-acuminate; staminode apices rounded or bifurcate.
→ 2
2. Staminode apices rounded or bifurcate; corollas 24–35 mm; stems glabrous or puberulent proximally.
var. alpinus
2. Staminode apices usually bifurcate; corollas 27–48 mm; stems puberulent to pubescent proximally.
var. brandegeei
Source FNA vol. 17, p. 167. FNA vol. 17, p. 137.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. glaber var. alpinus, P. glaber var. brandegeei, P. glaber var. glaber
Synonyms P. guadalupensis subsp. ernestii
Name authority Pursh: Fl. Amer. Sept. 2: 738. (1813) — (as Pentstemon glabra) A. Heller: Contr. Herb. Franklin Marshall Coll. 1: 92, plate 7. (1895) — (as Pentstemon)
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