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sawsepal penstemon, southern smooth beardtongue, western smooth beardtongue

sand dune penstemon, sand-dune beardtongue, sharp-leaf penstemon

Stems

ascending, (10–)50–65(–80) cm, glabrous or puberulent to pubescent, not glaucous.

ascending to erect, (9–)20–60 cm, glabrous or ± scabrous, especially distally.

Leaves

basal and cauline, or basal absent or reduced, not leathery, glabrous or puberulent to pubescent, not glaucous;

basal and proximal cauline 20–80(–155) × 5–20(–45) mm, blade obovate to oblanceolate or lanceolate, base tapered, margins entire, usually undulate, apex obtuse to acute, sometimes mucronate;

cauline (2 or)3–6(–8) pairs, sessile, 27–120(–150) × (6–)22–35(–43) mm, blade lanceolate, base truncate to cordate, apex obtuse to acute.

basal and cauline, glabrous, sometimes scabrous;

basal and proximal cauline (14–)30–95 × 4–15(–24) mm, blade oblanceolate to elliptic, base tapered, apex obtuse to acute or mucronate;

cauline (2 or)3–6(or 7) pairs, sessile, 18–60(–85) × (4–)8–28 mm, blade ovate to lanceolate, base tapered to cordate-clasping, apex acute to acuminate.

Thyrses

continuous, secund, (3–)6–26(–30) cm, axis glabrous or puberulent, verticillasters (3–)5–12, cymes 2–4-flowered, 2 per node;

proximal bracts ovate to lanceolate, 14–100 × 2–36 mm;

peduncles and pedicels glabrous or puberulent, peduncles to 71 mm, pedicels 4–14 mm.

continuous or interrupted, cylindric, 6–25(–30) cm, axis glabrous, rarely scabrous or glutinous, verticillasters (5–)7–15, cymes (2–)4–8(–12)-flowered;

proximal bracts ovate to lanceolate, 14–60 × 9–30 mm, margins entire, apex obtuse to acuminate or caudate;

peduncles and pedicels glabrous or scabrous, sometimes also glutinous, peduncles to 4 mm.

Flowers

calyx lobes ovate to orbiculate, 2–11 × 1.3–5.5 mm, glabrous or puberulent;

corolla blue to violet, with reddish purple nectar guides, ventricose, 24–48 mm, glabrous externally, glabrous internally or sparsely to moderately white-lanate internally abaxially, tube 5–12 mm, throat gradually inflated, slightly constricted at orifice, 8–13(–18) mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, pollen sacs divergent, navicular, 1.4–2.5(–2.8) mm, dehiscing incompletely, connective not splitting, sides sparsely to moderately hirsute, hairs white or tan, to 0.5 mm, rarely glabrous, sutures denticulate, teeth to 0.2 mm;

staminode (15–)18–22 mm, included or barely exserted, 0.6–1.3 mm diam., tip straight to recurved, rounded to distinctly bifurcate at apex, glabrous or distal 1–2 mm sparsely lanate, hairs pale yellow, to 1.5 mm;

style 18–20 mm.

calyx lobes lanceolate, (3.2–)4.5–10 × 1.2–3.2(–3.8) mm, margins entire or erose, herbaceous or narrowly scarious, glabrous or scabrous proximally and, sometimes, along margins, sometimes also glutinous;

corolla blue to violet or purple, with reddish purple nectar guides, tubular-funnelform, 11–20 mm, glabrous externally, glabrous internally, tube 5–9 mm, throat gradually inflated, 4–6 mm diam., rounded abaxially;

stamens included or longer pair reaching orifice to slightly exserted, pollen sacs opposite, navicular, 0.7–1.3(–1.5) mm, sutures papillate;

staminode 6–12 mm, reaching orifice, 0.6–1 mm diam., tip recurved, distal 1.5–3.5 mm densely pilose, hairs golden yellow, to 0.6 mm, hairs on proximal part of staminode mostly along margins;

style 8–15 mm.

Capsules

10–17 × 5–8 mm.

7–13 × 4–7 mm.

Penstemon glaber

Penstemon acuminatus

Distribution
from FNA
CO; MT; ND; NE; NM; SD; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
ID; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

The Penstemon glaber complex has been treated with as many as four species (F. W. Pennell 1920b) or one variable species as treated herein. M. K. W. Latady (1985) found clinal variation in most morphologic characters used to distinguish taxa in the complex and recognized only one species and two varieties: var. glaber (including var. alpinus) in the plains and mountains from southeastern Montana south to central Colorado; and var. brandegeei, in the mountains from central Colorado south to north-central New Mexico. A case for recognizing var. alpinus, as is done here, can be made on morphological, phenological, and ecological grounds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Penstemon acuminatus includes the northern var. acuminatus, which has denser thyrses than the southern var. latebracteatus. Herbage of var. acuminatus is usually scabrous; herbage of var. latebracteatus is rarely scabrous.

The Blackfoot tribe of the Central Rocky Mountains use Penstemon acuminatus as an analgesic, antiemetic, and gastrointestinal aid (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx lobes 2–4.8 mm, apices rounded or abruptly short-acuminate; staminode apices rounded.
var. glaber
1. Calyx lobes 3.8–11 mm, apices long-acuminate; staminode apices rounded or bifurcate.
→ 2
2. Staminode apices rounded or bifurcate; corollas 24–35 mm; stems glabrous or puberulent proximally.
var. alpinus
2. Staminode apices usually bifurcate; corollas 27–48 mm; stems puberulent to pubescent proximally.
var. brandegeei
1. Styles 11–15 mm; corollas 14–20 mm; pollen sacs 0.8–1.3(–1.5) mm; calyx lobes 5–10 mm.
var. acuminatus
1. Styles 8–11 mm; corollas 11–15 mm; pollen sacs 0.7–1 mm; calyx lobes (3.2–)4.5–7 mm.
var. latebracteatus
Source FNA vol. 17, p. 167. FNA vol. 17, p. 113.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. glaber var. alpinus, P. glaber var. brandegeei, P. glaber var. glaber
P. acuminatus var. acuminatus, P. acuminatus var. latebracteatus
Name authority Pursh: Fl. Amer. Sept. 2: 738. (1813) — (as Pentstemon glabra) Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1285. (1829) — (as Pentstemon acuminatum)
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