Gairdner's beardtongue, Gairdner's penstemon
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lance-leaf beardtongue
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Subshrubs. |
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ascending to erect, 6–40 cm, retrorsely hairy, not glaucous. |
24–75(–98) cm, sparsely to densely retrorsely hairy or puberulent, usually also glandular-pubescent distally. |
basal and cauline, alternate or mostly opposite, not leathery, retrorsely hairy; basal and proximal cauline 10–50(–70) × 1–4 mm, blade spatulate to oblanceolate or linear, base tapered, margins entire, revolute, apex rounded to obtuse or acute; cauline 8–12 pairs (or 13–25 leaves when alternate), sessile, 7–35 × 1–3 mm, blade oblanceolate to linear, base tapered, margins entire, apex acute. |
retrorsely hairy; basal and proximal cauline 8–80 × 3–12 mm, blade lanceolate or spatulate, base tapered, margins entire, apex rounded to obtuse or acute; cauline 7–15 pairs, 9–85(–110) × 1–11(–22) mm, blade lanceolate to linear, base tapered to truncate, apex acute to acuminate, rarely obtuse. |
interrupted, cylindric to secund, 2–12 cm, axis glandular-pubescent and retrorsely hairy, verticillasters 3–12, cymes 1(or 2)-flowered, 1 or 2 per node; proximal bracts lanceolate to linear, 5–18 × 0.5–3 mm, margins entire; peduncles and pedicels erect, glandular-pubescent and/or retrorsely hairy. |
± secund, 5–36 cm, axis glandular-pubescent, verticillasters 3–6, cymes 1 or 2(or 3)-flowered; proximal bracts lanceolate to linear, 5–16 × 1–3 mm; peduncles and pedicels glandular-pubescent, sometimes also retrorsely hairy. |
calyx lobes lanceolate, 5–8.5 × 1.7–2.7 mm, puberulent and glandular-pubescent; corolla violet to blue or purple, with or without reddish nectar guides, funnelform, 15–22 mm, glandular-pubescent externally, glandular-pubescent internally, tube 3–4 mm, throat gradually inflated, 5–6 mm diam., slightly 2-ridged abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, navicular to subexplanate, 0.8–1 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 8–11 mm, included, 0.4–0.6 mm diam., tip straight, distal 3–5 mm sparsely to densely pubescent, hairs yellow, to 1 mm; style 10–12 mm. |
calyx lobes ovate to lanceolate, 2.9–7 × 1.5–2.5(–3.6) mm, glandular-pubescent; corolla scarlet to red, unlined internally or lined on abaxial surface with faint reddish purple nectar guides, tubular-funnelform, 22–35 mm, glandular-pubescent internally abaxially, tube 5–6 mm, throat gradually inflated, 6–8 mm diam., rounded abaxially; stamens exserted (hidden by galeate adaxial lobes), pollen sacs parallel to divergent, 1.5–2 mm, sutures denticulate, teeth to 0.2 mm; staminode 15–17 mm, included, 0.3–0.4 mm diam.; style 25–30 mm, usually barely exserted from galea. |
5–9 × 3–5.3 mm, glabrous. |
9–14 × 6–7 mm. |
= 16. |
= 16. |
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Flowering May–Sep. |
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Gravelly pinyon-juniper woodlands, pine woodlands, thorn scrub, desert grasslands. |
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1200–1800 m. (3900–5900 ft.) |
ID; OR; WA
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AZ; NM; TX; Mexico (Aguascalientes, Chihuahua, Coahuila, Durango, Nayarit, Nuevo León, San Luis Potosí, Tamaulipas, Veracruz, Zacatecas) |
Varieties 2 (2 in the flora). Following A. Cronquist (1959), two varieties are recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon lanceolatus is known from southern Arizona (Cochise, Graham, and Greenlee counties), southern New Mexico (Doña Ana, Grant, Hidalgo, Luna, and Sierra counties), and western Texas (Brewster County). Crosswhite published Penstemon ramosus as an avowed substitute for the homonym P. pauciflorus Greene, distinguishing P. ramosus from P. lanceolatus by the former’s branched stems and relatively narrower leaves (1 mm wide versus 4–8 mm wide) with revolute margins. He mapped P. ramosus in southeastern Arizona and southwestern New Mexico and P. lanceolatus in western Texas and northern Mexico. Morphologic differences between these taxa are not consistent enough to warrant recognition of P. ramosus (J. L. Anderson et al. 2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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FNA vol. 17, p. 204. |
FNA vol. 17, p. 109. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Chamaeleon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. ramosus |
Hooker: Fl. Bor.-Amer. 2: 99. (1838) — (as Pentstemon) |
Bentham: Pl. Hartw., 22. (1839) — (as Pentstemon) |
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