Gairdner's beardtongue, Gairdner's penstemon
|
Deam's beardtongue
|
Subshrubs. |
Herbs. |
ascending to erect, 6–40 cm, retrorsely hairy, not glaucous. |
ascending to erect, 40–90 cm, retrorsely hairy proximally, usually retrorsely hairy distally, not glaucous. |
basal and cauline, alternate or mostly opposite, not leathery, retrorsely hairy; basal and proximal cauline 10–50(–70) × 1–4 mm, blade spatulate to oblanceolate or linear, base tapered, margins entire, revolute, apex rounded to obtuse or acute; cauline 8–12 pairs (or 13–25 leaves when alternate), sessile, 7–35 × 1–3 mm, blade oblanceolate to linear, base tapered, margins entire, apex acute. |
basal and cauline, basal sometimes withering by anthesis, not leathery, glabrate to puberulent; basal and proximal cauline 40–150 × 10–26 mm, blade oval to elliptic, base tapered, margins entire or ± denticulate, apex rounded to obtuse; cauline 7 or 8 pairs, sessile, 17–110 × 3–20 mm, blade oblanceolate to lanceolate, base tapered to clasping, margins entire or denticulate to serrulate, apex obtuse to acute. |
interrupted, cylindric to secund, 2–12 cm, axis glandular-pubescent and retrorsely hairy, verticillasters 3–12, cymes 1(or 2)-flowered, 1 or 2 per node; proximal bracts lanceolate to linear, 5–18 × 0.5–3 mm, margins entire; peduncles and pedicels erect, glandular-pubescent and/or retrorsely hairy. |
interrupted, conic, 11–22 cm, axis glabrous or sparsely glandular-pubescent, verticillasters 4–6, cymes (2–)5–8-flowered, 2 per node; proximal bracts lanceolate, 10–15 × 3–5 mm, margins entire; peduncles and pedicels ascending to erect, glabrous or sparsely glandular-pubescent. |
calyx lobes lanceolate, 5–8.5 × 1.7–2.7 mm, puberulent and glandular-pubescent; corolla violet to blue or purple, with or without reddish nectar guides, funnelform, 15–22 mm, glandular-pubescent externally, glandular-pubescent internally, tube 3–4 mm, throat gradually inflated, 5–6 mm diam., slightly 2-ridged abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, navicular to subexplanate, 0.8–1 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 8–11 mm, included, 0.4–0.6 mm diam., tip straight, distal 3–5 mm sparsely to densely pubescent, hairs yellow, to 1 mm; style 10–12 mm. |
calyx lobes ovate, 1.5–3.5(–4) × 0.9–1.5 mm, sparsely glandular-pubescent; corolla white, sometimes tinged lavender or purple, with reddish purple nectar guides, ventricose, 15–20(–22) mm, glandular-pubescent externally, sparsely white-pubescent internally abaxially, tube 5–6 mm, throat abruptly inflated, 5–7 mm diam., slightly 2-ridged abaxially; stamens included, pollen sacs opposite, navicular, 0.8–1 mm, dehiscing completely, connective splitting, sides glabrous, sometimes sparsely pubescent, hairs white, to 0.2 mm, sutures papillate; staminode 8–10 mm, reaching orifice, 0.3–0.4 mm diam., tip straight to slightly recurved, distal 3–4 mm pilose, hairs yellowish, to 1 mm; style 10–12 mm. |
5–9 × 3–5.3 mm, glabrous. |
5–8 × 4–5 mm, glabrous. |
= 16. |
|
|
|
|
|
|
Flowering May–Jun. |
|
Rocky oak woods, fields. |
|
100–200 m. (300–700 ft.) |
ID; OR; WA
|
IN |
Varieties 2 (2 in the flora). Following A. Cronquist (1959), two varieties are recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
A. C. Koelling (1964) assessed Penstemon deamii and its allies using morphological, cytological, and breeding studies. He concluded that P. deamii is a distinct member of the eastern North American polyploids that include P. calycosus, P. digitalis, and P. laevigatus. Geographically sympatric with P. calycosus and P. digitalis, P. deamii is distinguished from those two species by its shorter calyx lobes [1.5–3.5(–4) mm versus 4–9 mm], corollas [15–20(–22) mm versus (17–)20–35 mm], and pollen sacs (0.8–1 mm versus 1.2–1.7 mm). The species appears to be limited to Clark, Crawford, Floyd, Harrison, and Marion counties (K. Yatskievych 2000). Reports of P. deamii from southern Illinois appear to be based on specimens of questionable provenance or misidentifications (Koelling). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
|
FNA vol. 17, p. 204. |
FNA vol. 17, p. 199. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
|
|
Hooker: Fl. Bor.-Amer. 2: 99. (1838) — (as Pentstemon) |
Pennell: Monogr. Acad. Nat. Sci. Philadelphia 1: 212. (1935) |
| |