Fremont's beardtongue
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blowout beardtongue
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Plants: caudex rhizomelike. |
ascending to erect, 8–25(–40) cm, densely retrorsely hairy, not glaucous. |
decumbent to ascending, (15–)20–48 cm, glabrous. |
basal and cauline, or basal absent or reduced, not leathery, retrorsely hairy or glabrous, sometimes glabrescent, not glaucous; basal and proximal cauline 30–75(–100) × 2–18(–27) mm, blade oblanceolate to elliptic, base tapered, margins entire, apex obtuse to acute; cauline 2–4 pairs, sessile, rarely short-petiolate, 22–77 × 3–10(–15) mm, blade lanceolate, proximals sometimes oblanceolate, base tapered to truncate, apex acute. |
essentially cauline, basal absent or reduced, glabrous; basal and proximal cauline (25–)55–130(–175) × 3–25 mm, blade linear to lanceolate, base tapered, apex acute to acuminate; cauline 2–10 pairs, sessile, 60–110(–120) × 7–30 mm, blade lanceolate to linear, base clasping, apex acuminate to long-acuminate. |
interrupted to ± continuous, cylindric, (3–)8–22(–28) cm, axis moderately to densely retrorsely hairy, verticillasters (5–)7–15, cymes (1 or)2- or 3-flowered, 2 per node; proximal bracts lanceolate, 16–50(–63) × 3–10 mm; peduncles and pedicels retrorsely hairy. |
continuous, cylindric, (2–)6–21(–34) cm, axis glabrous, verticillasters (2–)6–10(–17), cymes 1–8-flowered; proximal bracts ovate, 58–120 × 20–45 mm; peduncles and pedicels glabrous. |
calyx lobes ovate, (3–)4–6.8 × 1.7–2.8 mm, glabrous, rarely ± puberulent; corolla blue to violet or purple, with violet nectar guides, funnelform, (12–)14–23(–28) mm, glabrous externally, glabrous internally, tube 6–8 mm, throat gradually inflated, not constricted at orifice, 6–9 mm diam., rounded abaxially; stamens included, pollen sacs divergent, navicular, (1–)1.2–1.5(–1.8) mm, dehiscing incompletely, proximal 1/4 indehiscent, connective not splitting, sides sparsely to moderately hispid, hairs white or tan, to 0.4(–1) mm, sutures papillate or denticulate, teeth to 0.1 mm; staminode 10–12 mm, included, 0.8–1.1 mm diam., tip recurved, distal 5–7 mm sparsely pilose, hairs yellow, to 1 mm; style 12–15 mm. |
calyx lobes lanceolate to linear, 8–13 × 1–3 mm, margins entire or erose, herbaceous or scarious, glabrous; corolla lavender to bluish, usually with magenta nectar guides, ampliate, 21–28 mm, glabrous externally, glabrous internally, tube 7–9 mm, throat abruptly inflated, 9–11 mm diam., rounded abaxially; stamens included, pollen sacs opposite, 1.1–2 mm, sutures papillate; staminode 13–16 mm, included, 1.2–3 mm diam., tip recurved, distal 0.1–5 mm sparsely to densely villous, hairs golden yellow, to 1 mm, rarely glabrous; style 13–23 mm. |
7–10 × 5–6 mm. |
8–12 × 5–9 mm. |
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= 16. |
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Flowering May–Jul(–Sep). |
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Blowouts in sand dunes. |
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900–2300 m. (3000–7500 ft.) |
CO; UT; WY
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NE; WY |
Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon haydenii is known from the Nebraska Sandhills, where extant populations occur in Box Butte, Cherry, Garden, Hooker, Morrill, and Sheridan counties. Historic populations occurred in Thomas County (D. M. Sutherland 1988). Penstemon haydenii was discovered in northern Carbon County, Wyoming, in the late 1990s some 300 km west of the nearest Nebraska populations; it might have been collected in Wyoming as early as 1877 (W. Fertig 2001). A morphometric analysis of Nebraska and Wyoming plants revealed differences that could justify recognition of Wyoming populations as a distinct variety (C. C. Freeman 2015). It is listed as endangered by the U.S. Department of the Interior. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Leaves densely retrorsely hairy, rarely glabrescent, basal and proximal cauline 2–18(–27) mm wide. | var. fremontii |
1. Leaves glabrous or glabrescent, except sometimes margins and midveins, basal and proximal cauline 2–12 mm wide. | var. glabrescens |
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FNA vol. 17, p. 166. |
FNA vol. 17, p. 119. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Torrey & A. Gray: Proc. Amer. Acad. Arts 6: 60. (1862) — (as Pentstemon fremonti) |
S. Watson: Bot. Gaz. 16: 311. (1891) — (as Pentstemon haydeni) |
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