Fremont's beardtongue
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California beardtongue, California penstemon
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ascending to erect, 8–25(–40) cm, densely retrorsely hairy, not glaucous. |
ascending to erect, (5–)8–30 cm, retrorsely hairy, hairs white, scalelike. |
basal and cauline, or basal absent or reduced, not leathery, retrorsely hairy or glabrous, sometimes glabrescent, not glaucous; basal and proximal cauline 30–75(–100) × 2–18(–27) mm, blade oblanceolate to elliptic, base tapered, margins entire, apex obtuse to acute; cauline 2–4 pairs, sessile, rarely short-petiolate, 22–77 × 3–10(–15) mm, blade lanceolate, proximals sometimes oblanceolate, base tapered to truncate, apex acute. |
± leathery or not, densely retrorsely hairy, hairs appressed, white, scalelike; cauline 10–30 pairs, petiolate, 8–16 × 1.5–2.5 mm, blade oblanceolate, base tapered, apex mucronate. |
interrupted to ± continuous, cylindric, (3–)8–22(–28) cm, axis moderately to densely retrorsely hairy, verticillasters (5–)7–15, cymes (1 or)2- or 3-flowered, 2 per node; proximal bracts lanceolate, 16–50(–63) × 3–10 mm; peduncles and pedicels retrorsely hairy. |
interrupted or continuous, secund, (1–)2–9 cm, axis retrorsely hairy with appressed, white, scalelike hairs, verticillasters 2–13, cymes 1–4-flowered, 1 per node; proximal bracts oblanceolate, 2–10 × 0.5–1.5 mm; peduncles and pedicels spreading to ascending, retrorsely hairy with appressed, white, scalelike hairs, and sparsely glandular-pubescent. |
calyx lobes ovate, (3–)4–6.8 × 1.7–2.8 mm, glabrous, rarely ± puberulent; corolla blue to violet or purple, with violet nectar guides, funnelform, (12–)14–23(–28) mm, glabrous externally, glabrous internally, tube 6–8 mm, throat gradually inflated, not constricted at orifice, 6–9 mm diam., rounded abaxially; stamens included, pollen sacs divergent, navicular, (1–)1.2–1.5(–1.8) mm, dehiscing incompletely, proximal 1/4 indehiscent, connective not splitting, sides sparsely to moderately hispid, hairs white or tan, to 0.4(–1) mm, sutures papillate or denticulate, teeth to 0.1 mm; staminode 10–12 mm, included, 0.8–1.1 mm diam., tip recurved, distal 5–7 mm sparsely pilose, hairs yellow, to 1 mm; style 12–15 mm. |
calyx lobes ovate, 3.5–5.5 × 1.5–2 mm, scarious-margined, retrorsely hairy with appressed, white, scalelike hairs, and sparsely glandular-pubescent; corolla purple to violet, lined internally abaxially with reddish purple nectar guides, tubular-funnelform, 14–18 mm, sparsely white-lanate internally abaxially, tube 5–6 mm, throat gradually inflated, 4.5–6 mm diam., 2-ridged abaxially; stamens included, filaments glabrous or sparsely puberulent and sometimes with appressed, white, scalelike hairs proximally, pollen sacs opposite, navicular, 0.9–1.4 mm, sutures papillate; staminode 7–9 mm, reaching orifice or exserted, flattened distally, 0.3–0.4 mm diam., tip straight to recurved, distal 4–5 mm pilose, hairs yellow, to 0.7 mm; style 9–11 mm. |
7–10 × 5–6 mm. |
5–7 × 4–5 mm. |
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Flowering May–Jul(–Aug). |
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Sandy or gravelly soils, pine-juniper woodlands, pine forests. |
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1200–2300 m. (3900–7500 ft.) |
CO; UT; WY
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CA; Mexico (Baja California)
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Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In California, Penstemon californicus is known from the San Jacinto Mountains in Orange and Riverside counties; it also occurs in the Sierra San Pedro Mártir in Baja California. The chromosome number of 2n = 16 reported by N. H. Holmgren (1993) is likely an error based on a count for Scrophularia californica reported in P. H. Raven et al. (1965) and later misattributed to P. californicus by D. V. Clark (1971) as n = 48 (the count listed in Raven et al.), and then corrected to 2n = 16. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Leaves densely retrorsely hairy, rarely glabrescent, basal and proximal cauline 2–18(–27) mm wide. | var. fremontii |
1. Leaves glabrous or glabrescent, except sometimes margins and midveins, basal and proximal cauline 2–12 mm wide. | var. glabrescens |
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FNA vol. 17, p. 166. |
FNA vol. 17, p. 101. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. linarioides var. californicus |
Torrey & A. Gray: Proc. Amer. Acad. Arts 6: 60. (1862) — (as Pentstemon fremonti) |
(Munz & I. M. Johnston) D. D. Keck: Bull. Torrey Bot. Club 64: 378. (1937) |
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