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Jone's penstemon, Jones' beardtongue

Habit Herbs, caulescent or essentially acaulescent (P. acaulis, P. yampaënsis).
Stems

prostrate, decumbent, ascending, or erect, 2–14(–20) cm, pubescent to densely retrorsely hairy.

retrorsely hairy, sometimes glandular-pubescent or glandular-villous distally or wholly, rarely glabrous, glabrate, or scabrous proximally or wholly, not glaucous.

Leaves

basal and cauline, not leathery, retrorsely hairy;

basal and proximal cauline petiolate, (5–)10–40(–55) × 4–14 mm, blade spatulate to oblanceolate, base tapered, margins entire, apex obtuse to acute, rarely rounded;

cauline 1–3 pairs, sessile, 12–38 × 3–7 mm, blade oblanceolate, not arcuate, base tapered, margins entire, apex obtuse to acute.

basal and cauline, sometimes ± cauline or ± basal, opposite, rarely whorled (P. albidus, P. auriberbis, P. jamesii), leathery or not, glabrous or glabrate, scabrous, retrorsely hairy, glandular-pubescent, pubescent, villous, or glandular-villous, not glaucous;

basal and proximal cauline petiolate, sometimes sessile, rarely short-petiolate (P. pumilus);

cauline sessile or short-petiolate, blade ovate, deltate-ovate, oblanceolate, lanceolate, elliptic, oblong, or linear, rarely pandurate, margins entire or toothed.

Thyrses

continuous, cylindric, 0.5–8(–10) cm, axis retrorsely hairy, verticillasters 1–4(–7), cymes 1- or 2(–4)-flowered, 1 or 2 per node;

proximal bracts oblanceolate to lanceolate, 12–34 × 2–8 mm;

peduncles and pedicels retrorsely hairy, sometimes sparsely glandular-pubescent.

continuous or interrupted, cylindric, rarely ± secund to secund, axis glandular-pubescent, glandular-villous, or retrorsely hairy, rarely puberulent or glabrous, cymes 1 or 2 per node;

peduncles and pedicels ascending to erect.

Flowers

calyx lobes lanceolate, 4–6 × 1–1.9 mm, retrorsely hairy, sometimes sparsely glandular-pubescent;

corolla light blue to blue or violet, with, rarely without, faint blue or reddish purple nectar guides, funnelform, 14–20 mm, glandular-pubescent externally, sparsely white-lanate internally abaxially, tube 6–7 mm, throat gradually inflated, not constricted at orifice, 4–7 mm diam., slightly 2-ridged abaxially;

stamens included or longer pair reaching orifice, pollen sacs opposite, navicular, 0.9–1.2 mm, dehiscing completely, sutures papillate;

staminode 7–10 mm, included or reaching orifice, 0.7–0.9 mm diam., tip recurved to coiled, distal 5–8 mm sparsely pubescent to lanate, hairs yellow or yellow-orange, to 1 mm;

style 10–12 mm.

calyx lobes: margins entire, sometimes erose (P. atwoodii, P. laricifolius, P. triflorus), herbaceous or ± scarious, glandular-pubescent or glandular-villous, sometimes also scabrous, rarely entirely retrorsely hairy, puberulent, or glabrous;

corolla white to lilac, lavender, blue, violet, purple, red, pink, or magenta, bilaterally symmetric, rarely nearly radially symmetric (P. goodrichii), weakly to strongly bilabiate, not personate, funnelform, tubular-funnelform, ventricose, or ampliate, glandular-pubescent externally, rarely glabrous (P. laricifolius), hairy internally abaxially, rarely glabrous or glandular-pubescent, throat gradually to abruptly inflated, rarely slightly inflated, constricted or not at orifice, rounded abaxially, sometimes slightly 2-ridged;

stamens included, reaching orifice, or longer pair exserted, filaments glabrous, pollen sacs parallel to divergent or opposite, navicular to subexplanate or explanate, rarely navicular-sigmoid, dehiscing completely, rarely incompletely, proximal 1/5–1/2 indehiscent, connective splitting, rarely not, sides glabrous, sutures smooth or papillate;

staminode included to exserted, flattened distally, 0.2–1.1 mm diam., tip straight to recurved or coiled, distal (40–)50–100% hairy, hairs to 4 mm;

style glabrous, sometimes proximally glandular-pubescent (P. eriantherus).

Capsules

4–6 × 4–5 mm.

glabrous, rarely glandular-pubescent distally.

Seeds

brown, dark brown, or black, angled to reniform or slightly rounded, 1.5–4.8 mm.

Penstemon dolius

Penstemon sect. Cristati

Phenology Flowering May–Jun.
Habitat Gravelly soils, shadscale and sagebrush shrublands, pinyon-juniper woodlands.
Elevation 1400–2100 m. (4600–6900 ft.)
Distribution
from FNA
NV; UT
[WildflowerSearch map]
[BONAP county map]
w North America; n Mexico
Discussion

Penstemon dolius is known from east-central Nevada (Elko, Nye, and White Pine counties) and west-central Utah (Beaver, Juab, Millard, Sanpete, and Tooele counties). Penstemon dolius is distinguished from P. duchesnensis by longer stems and leaves and more sparsely bearded staminodes. Corollas of P. dolius usually have nectar guides; P. duchesnensis appears to lack nectar guides.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 31 (31 in the flora).

Members of sect. Cristati are mostly calciphiles on soils derived from limestone, dolomite, or other calcareous substrates. Species in sect. Cristati generally have hairy herbage, glandular-pubescent inflorescences and flowers, and staminodes prominently bearded more than half their lengths. Two species often assigned elsewhere (Penstemon acaulis, P. laricifolius) are included in this section based on molecular data (A. D. Wolfe et al. 2006). D. D. Keck’s (1938) sect. Aurator, which appears widely in the literature, is invalid, published without a Latin diagnosis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems to 1 cm.
→ 2
2. Basal and proximal cauline leaves 6–15(–22) × 0.6–1.3(–1.5) mm, blades linear.
P. acaulis
2. Basal and proximal cauline leaves 15–25(–35) × 1.5–2.6(–4.5) mm, blades oblanceolate to linear.
P. yampaënsis
1. Stems 2–65(–100) cm.
→ 3
3. Corollas glabrous externally; calyx lobes glabrous.
P. laricifolius
3. Corollas glandular-pubescent externally; calyx lobes hairy.
→ 4
4. Pollen sacs parallel, navicular.
→ 5
5. Staminodes prominently exserted; basal and proximal cauline leaf margins sharply serrate-dentate, rarely entire; corollas ventricose.
P. pinorum
5. Staminodes included to barely exserted; basal and proximal cauline leaf margins entire or ± dentate; corollas funnelform or tubular-funnelform.
→ 6
6. Thyrses 8–30 cm, interrupted; stems (25–)30–60 cm; Arizona.
P. distans
6. Thyrses 2–7(–8) cm, continuous or interrupted; stems 2–25 cm; Utah.
→ 7
7. Leaves glabrous abaxially, retrorsely hairy adaxially, usually only along midveins on distal caulines; pollen sacs 1.3–1.6 mm.
P. franklinii
7. Leaves densely retrorsely hairy; pollen sacs 1.7–2 mm.
P. nanus
4. Pollen sacs opposite or divergent, explanate or subexplanate to navicular or navicular-sigmoid.
→ 8
8. Thyrse axes puberulent or retrorsely hairy.
→ 9
9. Leaves: basal and proximal cauline blades oblanceolate to linear, 1–4 mm wide, caulines usually arcuate; Idaho.
P. pumilus
9. Leaves: basal and proximal cauline blades spatulate to oblanceolate, 2–14 mm wide, caulines not arcuate; Nevada, Utah.
→ 10
10. Cauline leaves 12–38 × 3–7 mm; stems 2–14(–20) cm, prostrate, decumbent, ascending, or erect; corollas with, rarely without, faint blue or reddish purple nectar guides; ec Nevada, wc Utah.
P. dolius
10. Cauline leaves 14–23 × 1–4 mm; stems 2–6(–8) cm, ascending to erect; corollas without nectar guides; ne Utah.
P. duchesnensis
8. Thyrse axes glandular-pubescent or glandular-villous, sometimes retrorsely hairy.
→ 11
11. Capsules sparsely glandular-pubescent distally; styles sparsely glandular-pubescent proximally or glabrous.
P. eriantherus
11. Capsules glabrous; styles glabrous.
→ 12
12. Pollen sacs navicular or navicular-sigmoid.
→ 13
13. Corollas ventricose, 19–30 mm; staminodes prominently exserted.
P. gormanii
13. Corollas tubular-funnelform, 10–22(–24) mm; staminodes included to exserted.
→ 14
14. Stems puberulent, hairs usually white, scalelike; leaves densely retrorsely hairy, hairs white, scalelike; California.
P. monoensis
14. Stems glabrous or retrorsely hairy, sometimes sparsely glandular-pubescent distally, hairs not white or scalelike; leaves glabrous, puberulent, or retrorsely hairy, hairs not white or scalelike; Colorado, New Mexico, Utah.
→ 15
15. Pollen sacs divergent, dehiscing incompletely; corollas weakly bilabiate; Colorado, New Mexico.
P. auriberbis
15. Pollen sacs opposite, dehiscing completely; corollas bilabiate; Colorado, Utah.
→ 16
16. Basal and proximal cauline leaf blades oblanceolate to lanceolate; corollas 10–13 mm; styles 6–8 mm; Utah.
P. atwoodii
16. Basal and proximal cauline leaf blades spatulate to oblanceolate or ovate; corollas 15–20(–22) mm; styles 10–12 mm; Colorado, Utah.
P. moffatii
12. Pollen sacs explanate, rarely subexplanate.
→ 17
17. Corollas tubular-funnelform to funnelform.
→ 18
18. Corollas white, sometimes tinged pink or lavender.
→ 19
19. Leaves glabrate or puberulent to scabrous, basals and proximals (4–)7–18(–20) mm wide, blades oblanceolate or obovate to lanceolate; thyrse axes densely glandular-pubescent; Alberta, Manitoba, Ontario, Saskatchewan, Colorado, Iowa, Kansas, Minnesota, Montana, Nebraska, New Mexico, North Dakota, Oklahoma, South Dakota, Texas, Wyoming.
P. albidus
19. Leaves glabrous or glabrate, rarely basals retrorsely hairy along midveins, basals and proximals 1–6 mm wide, blades linear, rarely lanceolate; thyrse axes glandular-pubescent; Texas.
P. guadalupensis
18. Corollas blue, bluish lavender, lavender, pink, rose red, rose purple, violet, or purple.
→ 20
20. Corollas pink to rose red or rose purple; California, sw Nevada.
P. calcareus
20. Corollas lavender, violet, purple, bluish lavender, or blue; e Nevada, Utah.
→ 21
21. Staminodes exserted; corollas bilabiate, tubular-funnelform.
P. concinnus
21. Staminodes included; corollas weakly bilabiate, funnelform.
→ 22
22. Stems glabrous or slightly retrorsely hairy proximally, glandular-pubescent distally; basal and proximal cauline leaf blades lanceolate to linear; staminodes: distal 5–7 mm densely pilose, hairs yellow, to 0.8 mm.
P. goodrichii
22. Stems retrorsely hairy; basal and proximal cauline leaf blades oblanceolate to lanceolate; staminodes: distal 3–5 mm sparsely to densely pubescent, hairs yellow, to 1 mm.
P. marcusii
17. Corollas ventricose, ventricose-ampliate, or ampliate.
→ 23
23. Corollas 35–55 mm; staminodes 23–30 mm; styles 18–34 mm.
P. cobaea
23. Corollas 10–35 mm; staminodes 7–20 mm; styles 7–24 mm.
→ 24
24. Corollas glandular-pubescent internally abaxially, sometimes also pilose.
→ 25
25. Corollas 14–20(–22) mm; styles 11–13 mm; capsules 6–11 mm.
P. ophianthus
25. Corollas 22–35 mm; styles 17–24 mm; capsules (7–)10–16 mm.
→ 26
26. Staminodes exserted, distal 10–14 mm lanate, hairs to 3.5 mm; corolla throats abruptly inflated.
P. jamesii
26. Staminodes included or reaching orifice, distal 8–12 mm sparsely to moderately pilose, hairs to 1 mm; corolla throats gradually inflated.
P. triflorus
24. Corollas lanate or villous internally abaxially.
→ 27
27. Styles 15–24 mm; corollas 18–35 mm, tubes (6–)8–13 mm; staminodes 16–20 mm.
→ 28
28. Corollas 28–35 mm, ventricose-ampliate, sparsely white-lanate internally abaxially; Colorado, Utah.
P. grahamii
28. Corollas 18–28 mm, ventricose, densely white-lanate internally abaxially; California, Idaho, Nevada, Oregon.
P. janishiae
27. Styles 7–11 mm; corollas 10–22 mm, tubes 4–7 mm; staminodes 7–10 mm.
→ 29
29. Staminodes reaching orifices or exserted; corollas ampliate; leaves retrorsely hairy, sometimes glabrate; Arizona, Colorado, New Mexico, Utah.
P. breviculus
29. Staminodes exserted or prominently exserted; corollas ventricose; leaves retrorsely hairy, glabrate, or glabrous; California, Idaho, Nevada, Oregon.
→ 30
30. Corollas 10–14 mm; styles 7–9 mm; capsules 3–6 mm; California, Nevada.
P. barnebyi
30. Corollas 14–22 mm; styles 9–10 mm; capsules (6–)8–11 mm; Idaho, Oregon.
P. miser
Source FNA vol. 17, p. 133. FNA vol. 17, p. 125.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati Plantaginaceae > Penstemon > subg. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. acaulis, P. albidus, P. atwoodii, P. auriberbis, P. barnebyi, P. breviculus, P. calcareus, P. cobaea, P. concinnus, P. distans, P. dolius, P. duchesnensis, P. eriantherus, P. franklinii, P. goodrichii, P. gormanii, P. grahamii, P. guadalupensis, P. jamesii, P. janishiae, P. laricifolius, P. marcusii, P. miser, P. moffatii, P. monoensis, P. nanus, P. ophianthus, P. pinorum, P. pumilus, P. triflorus, P. yampaënsis
Synonyms P., P. section albidi
Name authority M. E. Jones ex Pennell: Contr. U.S. Natl. Herb. 20: 341. (1920) (Rydberg) Pennell: Contr. U.S. Natl. Herb. 20: 325, 328. (1920)
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