hot-rock beardtongue, hot-rock penstemon, rock penstemon, scabland penstemon
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pinto beardtongue, pinto penstemon, two-color beardtongue
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Subshrubs. |
Herbs. |
ascending to erect, (6–)15–50 cm, retrorsely hairy, sometime glabrate, not glaucous. |
ascending to erect, 60–150 cm, glaucous. |
cauline, opposite, subopposite, or whorled, not leathery, glabrous or retrorsely hairy; cauline 5–9 pairs, petiolate or sessile, (7–)10–50(–52) × (0.5–)1–20 mm, blade oblanceolate to ovate or spatulate, lanceolate to linear distally, base of proximals tapered, distals clasping, margins serrate to serrulate, rarely entire, apex rounded to obtuse, acute, or acuminate. |
basal and proximal cauline 37–110 × 10–50 mm, blade obovate, base tapered, margins coarsely serrate to coarsely dentate, apex obtuse to acute; cauline 4–8 pairs, sessile, 40–110 × 15–55 mm, blade ovate, base connate-perfoliate, margins coarsely serrate to coarsely dentate, apex acute to acuminate. |
interrupted, cylindric, 6–30 cm, axis ± glandular-pubescent, verticillasters 5–11, cymes 1–6-flowered, 2 per node; proximal bracts lanceolate to linear, 6–40 × 1–4 mm, margins entire or serrate to serrulate; peduncles and pedicels ascending to erect, glandular-pubescent. |
interrupted, secund, 16–90 cm, axis glandular-pubescent, verticillasters 9–23, cymes 1–4-flowered; proximal bracts ovate to lanceolate, 8–25 × 3–28 mm; peduncles and pedicels ascending to erect, glandular-pubescent. |
calyx lobes ovate to lanceolate, 2.3–7 × 0.6–1.9 mm, glandular-pubescent; corolla white to ochroleucous, sometimes tinged lavender, with brownish or purple nectar guides, funnelform, 9–18 mm, glabrate to glandular-pubescent externally, glandular-pubescent internally, sometimes glabrous or glabrate adaxially, tube 4–5 mm, throat slightly inflated, 2.5–5 mm diam., slightly 2-ridged abaxially; stamens: longer pair slightly exserted, pollen sacs opposite, explanate, 0.4–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 8–9 mm, exserted, 0.2–0.3 mm diam., tip straight, glabrous or distal 0.5–2.5 mm sparsely pilose or villous, hairs yellow or white, to 0.5 mm; style 7–10 mm. |
calyx lobes ovate, 4–6 × 2.2–3 mm, glandular-pubescent; corolla pink to rose pink, purple, or light yellow, with or without reddish or reddish purple nectar guides, strongly bilabiate, ventricose, 18–27 mm, glandular-pubescent externally, glandular-pubescent internally, and sparsely white- or yellowish lanate internally abaxially, tube 7–10 mm, length 1.7–2 times calyx lobes, throat abruptly inflated, constricted at orifice, 6–9 mm diam., rounded abaxially; stamens: longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen sacs explanate, 1.4–1.8 mm, sutures smooth; staminode 14–16 mm, included or exserted, 0.3–0.4 mm diam., tip recurved, distal 4–5 mm lanate, hairs yellow, to 2.8 mm, proximal 3–4 mm glandular-puberulent; style 14–16 mm, glabrous. |
3–5 × 2.5–4 mm, glabrous. |
10–13 × 5–7 mm, glabrous or sparsely glandular-pubescent distally. |
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Flowering Mar–Jun. |
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Gravelly soils, arroyos, roadsides, talus slopes, desert scrub, juniper woodlands. |
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500–1700 m. (1600–5600 ft.) |
CA; ID; MT; NV; OR; UT; WA; WY
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AZ; CA; NV
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Varieties 4 (4 in the flora). Penstemon deustus comprises four marginally discrete varieties. Extreme forms of each variety are distinctive, but extensive zones of contact exist among the varieties, and populations frequently exhibit degrees of intermediacy, especially in Oregon where all four varieties occur. The Paiute and Shoshoni tribes, centered in the Great Basin, used Penstemon deustus to treat dermatological, gastrointestinal, immunological, and other disorders (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon bicolor is known from the Mojave Desert in the Black Mountains of northwestern Mohave County, Arizona, the Castle, Clark, and New York mountains of eastern San Bernardino County, California, and the desert mountain ranges of southern and western Clark County, Nevada. Two subspecies differing in corolla color have been recognized, with yellow-corolla forms restricted to Clark County, Nevada. Subspecies are not recognized here, following J. D. Morefield (2006). Habitat destruction and genetic swamping caused by hybridization between Penstemon bicolor and P. palmeri threaten many populations of P. bicolor in the vicinity of Las Vegas, Nevada (G. Glenne 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Leaves whorled, sometimes opposite or subopposite, blades oblanceolate or lanceolate to linear, margins serrate to serrulate distally or entire. | var. variabilis |
1. Leaves opposite, blades ovate, spatulate, oblanceolate, or lanceolate, margins sharply to obscurely serrate, sometimes entire. | → 2 |
2. Calyx lobes lanceolate, (3.5–)4–7 mm, apices acuminate; corollas 9–12 mm; cauline leaf blade margins sharply serrate. | var. suffrutescens |
2. Calyx lobes ovate to lanceolate, 2.3–5(–6.3) mm, apices acute to acuminate; corollas 10–18 mm; cauline leaf blade margins obscurely to sharply serrate, sometimes entire. | → 3 |
3. Corollas 12–18 mm, glandular-pubescent externally, glandular-pubescent internally, sometimes obscurely adaxially, abaxial lobes white to ochroleucous, sometimes tinged lavender. | var. deustus |
3. Corollas 10–12(–15) mm, glabrate or glandular-pubescent externally, glabrous internally, or glabrate or glandular-pubescent abaxially and glabrous adaxially, abaxial lobes brown, sometimes white to ochroleucous. | var. pedicillatus |
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FNA vol. 17, p. 200. |
FNA vol. 17, p. 247. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. palmeri var. bicolor, P. bicolor subsp. roseus, P. pseudospectabilis subsp. bicolor |
Douglas ex Lindley: Edwards’s Bot. Reg. 16: plate 1318. (1830) — (as Pentstemon deustum) |
(Brandegee) Clokey & D. D. Keck: Bull. S. Calif. Acad. Sci. 38: 12. (1939) |
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