hot-rock beardtongue, hot-rock penstemon, rock penstemon, scabland penstemon
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Alamo beardtongue, los alamos beardtongue
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Subshrubs. |
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ascending to erect, (6–)15–50 cm, retrorsely hairy, sometime glabrate, not glaucous. |
ascending to erect, 14–120 cm, sometimes glaucous. |
cauline, opposite, subopposite, or whorled, not leathery, glabrous or retrorsely hairy; cauline 5–9 pairs, petiolate or sessile, (7–)10–50(–52) × (0.5–)1–20 mm, blade oblanceolate to ovate or spatulate, lanceolate to linear distally, base of proximals tapered, distals clasping, margins serrate to serrulate, rarely entire, apex rounded to obtuse, acute, or acuminate. |
not leathery, glabrous, sometimes glaucous; basal and proximal cauline 25–140 × 9–45 mm, blade elliptic to obovate or lanceolate, base tapered, margins entire, apex obtuse to acute; cauline 2–6 pairs, sessile or proximals short-petiolate, 10–135 × 3–30 mm, blade oblong to oblanceolate or lanceolate, base tapered to truncate, margins entire, apex obtuse to acute. |
interrupted, cylindric, 6–30 cm, axis ± glandular-pubescent, verticillasters 5–11, cymes 1–6-flowered, 2 per node; proximal bracts lanceolate to linear, 6–40 × 1–4 mm, margins entire or serrate to serrulate; peduncles and pedicels ascending to erect, glandular-pubescent. |
interrupted, cylindric to ± secund, 12–34(–90) cm, axis glabrous, verticillasters 5–15(–30), cymes (1 or)2(–4)-flowered; proximal bracts lanceolate to subulate, 4–15 × 1–5 mm; peduncles and pedicels ascending to erect, peduncles glabrous, pedicels glabrous or sparsely glandular-pubescent. |
calyx lobes ovate to lanceolate, 2.3–7 × 0.6–1.9 mm, glandular-pubescent; corolla white to ochroleucous, sometimes tinged lavender, with brownish or purple nectar guides, funnelform, 9–18 mm, glabrate to glandular-pubescent externally, glandular-pubescent internally, sometimes glabrous or glabrate adaxially, tube 4–5 mm, throat slightly inflated, 2.5–5 mm diam., slightly 2-ridged abaxially; stamens: longer pair slightly exserted, pollen sacs opposite, explanate, 0.4–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 8–9 mm, exserted, 0.2–0.3 mm diam., tip straight, glabrous or distal 0.5–2.5 mm sparsely pilose or villous, hairs yellow or white, to 0.5 mm; style 7–10 mm. |
calyx lobes lanceolate, 3–4.5 × 1–1.5 mm, margins entire, sparsely glandular-pubescent; corolla red to orangish red, without nectar guides, nearly radially symmetric, weakly bilabiate, tubular-funnelform, 19–25 mm, glandular-pubescent externally, glandular-pubescent internally abaxially, tube 6–8 mm, throat gradually inflated, 5–7 mm diam., rounded abaxially; stamens included, pollen sacs explanate, 1–1.4 mm, sutures smooth; staminode 7–9 mm, flattened distally, 0.3–0.5 mm diam., tip straight to recurved, glabrous; style 9–11 mm. |
3–5 × 2.5–4 mm, glabrous. |
11–16 × 5–8 mm. |
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= 16. |
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Flowering Apr–Jun. |
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Limestone canyons, cliffs, and slopes. |
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1300–1600 m. (4300–5200 ft.) |
CA; ID; MT; NV; OR; UT; WA; WY
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NM; TX |
Varieties 4 (4 in the flora). Penstemon deustus comprises four marginally discrete varieties. Extreme forms of each variety are distinctive, but extensive zones of contact exist among the varieties, and populations frequently exhibit degrees of intermediacy, especially in Oregon where all four varieties occur. The Paiute and Shoshoni tribes, centered in the Great Basin, used Penstemon deustus to treat dermatological, gastrointestinal, immunological, and other disorders (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon alamosensis is known from the Sacramento and San Andres mountains in New Mexico (Doña Ana, Lincoln, and Otero counties) and in the Hueco Mountains in western Texas (El Paso County). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Leaves whorled, sometimes opposite or subopposite, blades oblanceolate or lanceolate to linear, margins serrate to serrulate distally or entire. | var. variabilis |
1. Leaves opposite, blades ovate, spatulate, oblanceolate, or lanceolate, margins sharply to obscurely serrate, sometimes entire. | → 2 |
2. Calyx lobes lanceolate, (3.5–)4–7 mm, apices acuminate; corollas 9–12 mm; cauline leaf blade margins sharply serrate. | var. suffrutescens |
2. Calyx lobes ovate to lanceolate, 2.3–5(–6.3) mm, apices acute to acuminate; corollas 10–18 mm; cauline leaf blade margins obscurely to sharply serrate, sometimes entire. | → 3 |
3. Corollas 12–18 mm, glandular-pubescent externally, glandular-pubescent internally, sometimes obscurely adaxially, abaxial lobes white to ochroleucous, sometimes tinged lavender. | var. deustus |
3. Corollas 10–12(–15) mm, glabrate or glandular-pubescent externally, glabrous internally, or glabrate or glandular-pubescent abaxially and glabrous adaxially, abaxial lobes brown, sometimes white to ochroleucous. | var. pedicillatus |
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FNA vol. 17, p. 200. |
FNA vol. 17, p. 151. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Douglas ex Lindley: Edwards’s Bot. Reg. 16: plate 1318. (1830) — (as Pentstemon deustum) |
Pennell & G. T. Nisbet: Univ. Kansas Sci. Bull. 41: 709, figs. 41, 43, 45. (1960) |
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