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blue-stem beardtongue, bluestem penstemon

Platy or red canyon beardtongue, red canyon beardtongue

Habit Plants: caudex rhizomelike.
Stems

ascending to erect, (16–)20–60 cm, glabrous or ± puberulent proximally, ± glaucous.

decumbent to erect, 3–10(–15) cm, glabrous.

Leaves

basal and cauline, ± leathery or not, glabrous or proximals puberulent proximally or along midveins, ± glaucous;

basal and proximal cauline (30–)50–95 × 10–15(–28) mm, blade spatulate to oblanceolate, base tapered, margins entire, usually undulate, apex rounded to obtuse;

cauline 2–4 pairs, sessile, 20–73 × 4–15 mm, blade lanceolate to elliptic, base truncate to clasping, margins crisped or undulate, apex acute.

basal and cauline, glabrous;

basal and proximal cauline (10–)20–35 × 5–17 mm, blade obovate, base tapered, apex rounded to obtuse;

cauline 1–3 pairs, sessile, 10–29 × 5–12 mm, blade ovate to elliptic, base tapered, apex rounded to mucronate.

Thyrses

interrupted, secund to subsecund, (4–)8–31 cm, axis glabrous, verticillasters 5–9(–12), cymes (1 or)2–5-flowered, 2 per node;

proximal bracts ovate to lanceolate, 15–50 × 3–23 mm;

peduncles and pedicels glabrous, peduncles to 18 mm, pedicels 1–7 mm.

continuous, cylindric, 1–6 cm, axis glabrous, verticillasters 1–5(or 6), cymes 3–5-flowered;

proximal bracts ovate, 6–14 × 3–13 mm;

peduncles and pedicels glabrous.

Flowers

calyx lobes ovate, 3.5–4.5 × 1.4–2 mm, apex acute to acuminate, glabrous;

corolla blue to violet, with violet nectar guides, funnelform to weakly ventricose, 14–18(–20) mm, glabrous or ± glandular externally, glabrous internally, tube 5–8 mm, throat gradually inflated, not constricted at orifice, 5–7 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, pollen sacs divergent, navicular-sigmoid, 1.2–1.7 mm, dehiscing incompletely, proximal 1/4 indehiscent, connective not splitting, sides sparsely to moderately pilose, hairs white, to 0.5(–0.7) mm, sutures denticulate, teeth to 0.1 mm;

staminode: 9–11 mm, reaching orifice, 0.2–0.4 mm diam., tip straight, distal 2–6 mm sparsely pilose, hairs golden yellow, to 0.6 mm;

style 9–12 mm.

calyx lobes ovate to lanceolate, (3–)4–5.5 × 1.7–2.2 mm, margins entire or erose, narrowly to broadly scarious, glabrous;

corolla blue to violet or lavender, with reddish purple nectar guides, tubular-funnelform, 13–17(–19) mm, glabrous externally, sparsely white-villous internally abaxially, tube 4–5 mm, throat gradually inflated, 4–5 mm diam., rounded abaxially;

stamens included, pollen sacs divergent to opposite, 0.8–1.2 mm, sutures smooth or obscurely papillate;

staminode 6–8 mm, included, 0.8–1 mm diam., tip recurved, distal 4–5 mm moderately to densely pubescent, hairs golden yellow, to 1 mm;

style 7–8 mm.

Capsules

6–10 × 4–5 mm.

5–7 × 5–6 mm.

Penstemon cyanocaulis

Penstemon bracteatus

Phenology Flowering Apr–Jun. Flowering May–Jul.
Habitat Rocky slopes, pinyon-juniper, oak, and oak-juniper woodlands. Gravelly and talus slopes, pine woodlands.
Elevation 1300–2700 m. (4300–8900 ft.) 2100–2500 m. (6900–8200 ft.)
Distribution
from FNA
CO; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon cyanocaulis is known from Mesa, Montrose, and San Miguel counties, Colorado, and Carbon, Emery, Grand, and San Juan counties, Utah. It has crisped or undulate leaf margins that help distinguish it in the field.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon bracteatus is known from Garfield, Iron, and Kane counties. Plants grow on limestones of the Claron Formation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 165. FNA vol. 17, p. 116.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority Payson: Bot. Gaz. 60: 380. (1915) — (as Pentstemon) D. D. Keck: Leafl. W. Bot. 1: 82. (1934)
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