Cusick's beardtongue, Cusick's penstemon
|
Arkansas beardtongue
|
Herbs or subshrubs. |
Herbs. |
ascending to erect, 15–45 cm, retrorsely hairy, hairs pointed, not glaucous. |
ascending to erect, (18–)25–45(–61) cm, retrorsely hairy, sometimes also ± glandular-pubescent distally, not glaucous. |
cauline, opposite or subalternate, retrorsely hairy, hairs pointed, not glaucous; cauline 4–8 pairs, short-petiolate or sessile, 16–70 × 1–8(–14) mm, blade lanceolate to linear, base tapered, margins entire, apex obtuse to acute. |
basal and cauline, basal often withering by anthesis, not leathery, glabrous or sparsely retrorsely hairy, especially along midvein and margins, abaxially, glabrous or sparsely puberulent adaxially; basal and proximal cauline 20–95 × 5–28 mm, blade spatulate or obovate to ovate, base tapered, margins subentire to ± serrate or dentate, apex rounded to obtuse or acute; cauline (6–)8–12 pairs, short-petiolate or sessile, 22–80 × 4–20 mm, blade lanceolate, base tapered to clasping, margins ± serrate or dentate, apex acute to acuminate. |
interrupted, cylindric, 4–15 cm, axis puberulent or retrorsely hairy, verticillasters 3–10, cymes 1–5(–7)-flowered, 2 per node; proximal bracts lanceolate to linear, 7–50 × 1–6 mm; peduncles and pedicels ascending to erect, puberulent or retrorsely hairy. |
interrupted, conic, 6–24 cm, axis retrorsely hairy and ± glandular-pubescent, verticillasters 4–8, cymes 3–13-flowered, 2 per node; proximal bracts lanceolate, 6–40 × 2–12 mm, margins entire or ± dentate; peduncles and pedicels ascending, retrorsely hairy and ± glandular-pubescent. |
calyx lobes ovate, 3–5.5 × 1.4–2.3 mm, glabrous or sparsely puberulent; corolla blue to violet or purple, without nectar guides, ampliate, 16–21 mm, glabrous externally, glabrous internally, tube 4–5 mm, throat gradually inflated, 5–6 mm diam., slightly 2-ridged abaxially; stamens included, filaments glabrous, pollen sacs parallel, 1–1.5(–1.8) mm, distal 1/2–2/3 indehiscent, sides hispidulous, hairs white, to 0.1 mm near filament attachment, sutures denticulate, teeth to 0.1 mm; staminode 10–12 mm, reaching orifice or barely exserted, 0.5–0.7 mm diam., glabrous; style 11–13 mm. |
calyx lobes ovate, 2.8–4.5 × 1.1–2 mm, sparsely glandular-pubescent; corolla white, with reddish purple nectar guides, tubular to tubular-funnelform, 15–19 mm, glandular-pubescent externally, moderately whitish lanate internally abaxially, tube 4–6 mm, throat slightly inflated, 4–7 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular, 0.8–1 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 10–12 mm, exserted, 0.2–0.4 mm diam., tip straight to slightly recurved, distal 5–7 mm ± villous, hairs yellow or golden yellow, to 1.2 mm; style 9–11 mm. |
6–7 × 4.5–6 mm. |
4–7 × 4–5 mm, glabrous. |
= 16. |
= 16. |
|
|
|
|
Flowering May–Jul. |
Flowering Apr–Jun. |
Rocky, often basaltic, sagebrush shrublands. |
Rocky oak-hickory woodlands, glades, rocky stream bottoms. |
600–1500 m. (2000–4900 ft.) |
60–300 m. (200–1000 ft.) |
ID; OR
|
AR; MO; OK
|
Penstemon cusickii is known from west-central and southwestern Idaho (Ada, Blaine, Canyon, Elmore, Gem, Owyhee, and Washington counties) and eastern Oregon (Baker, Grant, Harney, Malheur, Union, and Wheeler counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon arkansanus is centered in the Interior Highlands. The species is most similar to P. pallidus, from which it differs by its narrower, less hairy, and more numerous cauline leaves. Some specimens from the narrow zone of sympatry in north-central Arkansas and south-central Missouri exhibit varying degrees of morphologic intermediacy. Penstemon arkansanus has been reported from Illinois (R. H. Mohlenbrock 1986) and Texas (S. D. Jones et al. 1997); those reports seem unlikely. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 232. |
FNA vol. 17, p. 193. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
|
P. multicaulis, P. pallidus subsp. arkansanus, P. wherryi |
A. Gray: Proc. Amer. Acad. Arts 16: 106. (1880) — (as Pentstemon) |
Pennell: Proc. Acad. Nat. Sci. Philadelphia 73: 493. (1922) |
| |