Crandall's beardtongue
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pine-leaf beardtongue, pineneedle beardtongue, pineneedle penstemon
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prostrate, decumbent, ascending, or erect, 2–28 cm, retrorsely hairy or pubescent. |
erect, (5–)10–50 cm, glabrous or retrorsely hairy, hairs pointed. |
not leathery, glabrous, glabrate, scabrous, or pubescent, hairs pointed; cauline (1 or)2–4 pairs, petiolate, (2–)4–47 × 0.5–8 mm, blade elliptic to obovate, oblanceolate, or linear, base tapered, apex acute to mucronate. |
leathery, sparsely retrorsely hairy, hairs pointed, especially along margins, or glabrous; cauline 10–30 pairs, sessile, 4–32 × 0.5–1.2 mm, blade linear, base tapered to clasping, apex acuminate. |
continuous, ± secund, 1–8(–16) cm, axis retrorsely hairy, hairs pointed and, usually, sparsely glandular-pubescent, verticillasters (1 or)2–8, cymes 1–4-flowered, 1 or 2 per node; proximal bracts oblanceolate to linear, 10–25 × 0.5–4 mm; peduncles and pedicels ascending to erect, retrorsely hairy and, usually, sparsely glandular-pubescent. |
interrupted, secund, (1–)3–15 cm, axis retrorsely hairy to puberulent, or glabrous, verticillasters 1–7, cymes 1–3-flowered, 1(or 2) per node; proximal bracts linear, 3–14 × 0.3–1 mm; peduncles and pedicels ascending to erect, glandular-pubescent, sometimes puberulent. |
calyx lobes lanceolate, 4–7(–7.5) × 0.9–1.8(–2.2) mm, broadly scarious-margined, apex acuminate to caudate, retrorsely puberulent, hairs pointed, and sparsely glandular-pubescent; corolla blue to violet or lavender, lined internally abaxially with violet or reddish purple nectar guides, funnelform, 14–20 mm, sparsely to densely yellow-lanate or yellow-pilose internally abaxially, tube 6–8 mm, throat gradually inflated, 6–8 mm diam., 2-ridged abaxially; stamens: longer pair reaching orifice, filaments glabrous, pollen sacs opposite, navicular to subexplanate, 0.9–1.4 mm, sutures papillate; staminode 11–14 mm, reaching orifice or exserted, flattened distally, 0.3–0.4 mm diam., tip straight to recurved, distal 7–10 mm densely pilose, hairs golden yellow, to 1 mm; style 13–15 mm. |
calyx lobes lanceolate, 5–7 × 1.5–2 mm, scarious-margined, glandular-pubescent, hairs pointed; corolla scarlet, throat sometimes yellow or yellow- or orange-spotted, unlined internally, salverform, 25–32 mm, yellow-lanate internally abaxially, tube 4–6 mm, throat slightly inflated, 4–5 mm diam., slightly 2-ridged abaxially; stamens exserted (often hidden by galeate adaxial lobes), filaments glabrous, pollen sacs opposite, explanate, 0.8–1 mm, sutures smooth; staminode 12–17 mm, included, slightly flattened distally, 0.1–0.2 mm diam., tip straight, distal 9–10 mm densely pilose, hairs yellow, to 1.3 mm; style 21–27 mm. |
7–8 × 3.5–4.5 mm. |
5–9 × 3–4 mm. |
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= 16. |
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Flowering Jun–Jul(–Sep). |
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Rocky slopes, cliffs. |
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1800–3100 m. (5900–10200 ft.) |
CO; NM; UT
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AZ; NM; Mexico (Chihuahua, Sonora)
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Varieties 5 (5 in the flora). Penstemon crandallii comprises five partly sympatric, sometimes intergrading varieties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon pinifolius has been included as an anomalous member of sect. Leptostemon Trautvetter; R. M. Straw (1962) expressed doubt about that association. It was allied with members of sect. Caespitosi in the molecular study by A. D. Wolfe et al. (2006). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Leaves glabrous or sparsely puberulent, especially along margins, blades obovate to elliptic; stems prostrate or decumbent; Colorado. | var. procumbens |
1. Leaves glabrate or scabrous to pubescent, blades linear, oblanceolate, or obovate; stems decumbent, ascending, or erect; Colorado, New Mexico, Utah. | → 2 |
2. Leaf blades obovate or oblanceolate; stems decumbent to ascending. | → 3 |
3. Leaves glabrate or scabrous, blades oblanceolate; Colorado, Utah. | var. crandallii |
3. Leaves ± scabrous, blades obovate or oblanceolate; Utah. | var. atratus |
2. Leaf blades linear to oblanceolate; stems ascending to erect. | → 4 |
4. Stems retrorsely hairy or puberulent, ascending to erect; Colorado, New Mexico. | var. glabrescens |
4. Stems puberulent to pubescent, erect; Colorado. | var. ramaleyi |
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FNA vol. 17, p. 101. |
FNA vol. 17, p. 105. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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A. Nelson: Bull. Torrey Bot. Club 26: 354. (1899) — (as Pentstemon) |
Greene: Bot. Gaz. 6: 218. (1881) — (as Pentstemon) |
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