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Crandall's beardtongue

Berry's penstemon, mountain pride, Newberry's beardtongue

Habit Subshrubs, cespitose.
Stems

prostrate, decumbent, ascending, or erect, 2–28 cm, retrorsely hairy or pubescent.

ascending to erect, (5–)10–32 cm, glabrous or retrorsely hairy, not glaucous.

Leaves

not leathery, glabrous, glabrate, scabrous, or pubescent, hairs pointed;

cauline (1 or)2–4 pairs, petiolate, (2–)4–47 × 0.5–8 mm, blade elliptic to obovate, oblanceolate, or linear, base tapered, apex acute to mucronate.

persistent, 4–9 pairs, distals distinctly to slightly smaller than proximals, short-petiolate or sessile, (5–)9–42 × 4–14(–18) mm, blade elliptic to obovate, base tapered, cuneate, or cordate-clasping, margins ± serrulate or serrulate-denticulate to serrate, apex rounded to obtuse or acute, glabrous, glaucous.

Thyrses

continuous, ± secund, 1–8(–16) cm, axis retrorsely hairy, hairs pointed and, usually, sparsely glandular-pubescent, verticillasters (1 or)2–8, cymes 1–4-flowered, 1 or 2 per node;

proximal bracts oblanceolate to linear, 10–25 × 0.5–4 mm;

peduncles and pedicels ascending to erect, retrorsely hairy and, usually, sparsely glandular-pubescent.

continuous, ± secund, 2–20 cm, axis glandular-pubescent, verticillasters 4–12, cymes 1-flowered;

proximal bracts ovate to lanceolate, 4–11(–20) × 2–8(–15) mm;

peduncles and pedicels ascending, glandular-pubescent.

Flowers

calyx lobes lanceolate, 4–7(–7.5) × 0.9–1.8(–2.2) mm, broadly scarious-margined, apex acuminate to caudate, retrorsely puberulent, hairs pointed, and sparsely glandular-pubescent;

corolla blue to violet or lavender, lined internally abaxially with violet or reddish purple nectar guides, funnelform, 14–20 mm, sparsely to densely yellow-lanate or yellow-pilose internally abaxially, tube 6–8 mm, throat gradually inflated, 6–8 mm diam., 2-ridged abaxially;

stamens: longer pair reaching orifice, filaments glabrous, pollen sacs opposite, navicular to subexplanate, 0.9–1.4 mm, sutures papillate;

staminode 11–14 mm, reaching orifice or exserted, flattened distally, 0.3–0.4 mm diam., tip straight to recurved, distal 7–10 mm densely pilose, hairs golden yellow, to 1 mm;

style 13–15 mm.

calyx lobes lanceolate, 6–10 × 2–2.5 mm, glandular-pubescent;

corolla pink, red, scarlet, lavender, or purple, essentially unlined internally but abaxial ridges sometimes pink to whitish, not personate, funnelform, 22–35 mm, glabrous externally, moderately to densely white-lanate internally abaxially, tube 6–9 mm, throat 5–12 mm diam.;

stamens included or longer pair exserted, pollen sacs 0.8–1.7 mm;

staminode 8–14 mm, slightly flattened distally, 0.1–0.3 mm diam., tip straight, distal 2–8 mm pubescent to lanate, hairs white or yellowish, to 1 mm;

style 20–24 mm.

Capsules

7–8 × 3.5–4.5 mm.

(8–)10–13 × (4–)5–8 mm.

Penstemon crandallii

Penstemon newberryi

Distribution
from FNA
CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 5 (5 in the flora).

Penstemon crandallii comprises five partly sympatric, sometimes intergrading varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Penstemon newberryi is known from the Sierra Nevada and Cascade ranges of California, Nevada, and Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves glabrous or sparsely puberulent, especially along margins, blades obovate to elliptic; stems prostrate or decumbent; Colorado.
var. procumbens
1. Leaves glabrate or scabrous to pubescent, blades linear, oblanceolate, or obovate; stems decumbent, ascending, or erect; Colorado, New Mexico, Utah.
→ 2
2. Leaf blades obovate or oblanceolate; stems decumbent to ascending.
→ 3
3. Leaves glabrate or scabrous, blades oblanceolate; Colorado, Utah.
var. crandallii
3. Leaves ± scabrous, blades obovate or oblanceolate; Utah.
var. atratus
2. Leaf blades linear to oblanceolate; stems ascending to erect.
→ 4
4. Stems retrorsely hairy or puberulent, ascending to erect; Colorado, New Mexico.
var. glabrescens
4. Stems puberulent to pubescent, erect; Colorado.
var. ramaleyi
1. Corollas pink to lavender or purple, 27–35 mm; stamens included, pollen sacs 1.2–1.7 mm.
var. berryi
1. Corollas red, scarlet, or purple, 22–30 mm; stamens: longer pair exserted, pollen sacs 0.8–1.3 mm.
→ 2
2. Corollas red to scarlet; distal leaves distinctly smaller than proximals.
var. newberryi
2. Corollas purple; distal leaves slightly smaller than proximals.
var. sonomensis
Source FNA vol. 17, p. 101. FNA vol. 17, p. 90.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi Plantaginaceae > Penstemon > subg. Dasanthera > sect. Erianthera
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. crandallii var. atratus, P. crandallii var. crandallii, P. crandallii var. glabrescens, P. crandallii var. procumbens, P. crandallii var. ramaleyi
P. newberryi var. berryi, P. newberryi var. newberryi, P. newberryi var. sonomensis
Name authority A. Nelson: Bull. Torrey Bot. Club 26: 354. (1899) — (as Pentstemon) A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 6(3): 82, plate 14. (1858) — (as Pentstemon)
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