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Crandall's beardtongue

bunchleaf penstemon, foothill beardtongue, foothill penstemon

Habit Subshrubs.
Stems

prostrate, decumbent, ascending, or erect, 2–28 cm, retrorsely hairy or pubescent.

ascending to erect, 30–150 cm, glabrous or retrorsely hairy, hairs pointed, rarely glaucous.

Leaves

not leathery, glabrous, glabrate, scabrous, or pubescent, hairs pointed;

cauline (1 or)2–4 pairs, petiolate, (2–)4–47 × 0.5–8 mm, blade elliptic to obovate, oblanceolate, or linear, base tapered, apex acute to mucronate.

essentially cauline, basal absent or essentially so, opposite, glabrous or retrorsely hairy to puberulent, hairs pointed, rarely glaucous;

cauline 5–11 pairs, petiolate or sessile, 10–100 × 0.5–12 mm, blade oblanceolate to lanceolate or linear, base tapered to clasping, margins entire, apex acute.

Thyrses

continuous, ± secund, 1–8(–16) cm, axis retrorsely hairy, hairs pointed and, usually, sparsely glandular-pubescent, verticillasters (1 or)2–8, cymes 1–4-flowered, 1 or 2 per node;

proximal bracts oblanceolate to linear, 10–25 × 0.5–4 mm;

peduncles and pedicels ascending to erect, retrorsely hairy and, usually, sparsely glandular-pubescent.

interrupted or continuous, cylindric, (6–)10–32 cm, axis glabrous or retrorsely hairy, verticillasters (3–)5–14, cymes 1(–3)-flowered, (1 or)2 per node;

proximal bracts lanceolate to linear, 5–65 × 0.5–5 mm;

peduncles and pedicels ascending to erect, glabrous or retrorsely hairy.

Flowers

calyx lobes lanceolate, 4–7(–7.5) × 0.9–1.8(–2.2) mm, broadly scarious-margined, apex acuminate to caudate, retrorsely puberulent, hairs pointed, and sparsely glandular-pubescent;

corolla blue to violet or lavender, lined internally abaxially with violet or reddish purple nectar guides, funnelform, 14–20 mm, sparsely to densely yellow-lanate or yellow-pilose internally abaxially, tube 6–8 mm, throat gradually inflated, 6–8 mm diam., 2-ridged abaxially;

stamens: longer pair reaching orifice, filaments glabrous, pollen sacs opposite, navicular to subexplanate, 0.9–1.4 mm, sutures papillate;

staminode 11–14 mm, reaching orifice or exserted, flattened distally, 0.3–0.4 mm diam., tip straight to recurved, distal 7–10 mm densely pilose, hairs golden yellow, to 1 mm;

style 13–15 mm.

calyx lobes obovate to oblanceolate or ovate to lanceolate, 2.6–6(–7) × 1.3–3.3 mm, glabrous or puberulent;

corolla blue to violet or purple, without nectar guides, ventricose to ventricose-ampliate, 24–38 mm, glabrous externally, glabrous internally, tube 7–10 mm, throat gradually inflated, 8–11 mm diam., 2-ridged abaxially;

stamens included or longer pair reaching orifice or slightly exserted, filaments glabrous, pollen sacs parallel, 2.2–3 mm, distal 1/3–1/2 indehiscent, sides pubescent to lanate, hairs white, to 0.8 mm near filament attachment, sutures denticulate, teeth to 0.2 mm;

staminode 16–19 mm, included, 0.4–0.6 mm diam., glabrous;

style 18–27 mm.

Capsules

7–8 × 3.5–4.5 mm.

6–9 × 4–5 mm.

2n

= 16, 32.

Penstemon crandallii

Penstemon heterophyllus

Distribution
from FNA
CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 5 (5 in the flora).

Penstemon crandallii comprises five partly sympatric, sometimes intergrading varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Penstemon heterophyllus apparently occurs only in California (see discussion under 209c. var. purdyi). The species includes three varieties concentrated in the Coast Ranges from Humboldt County south to San Bernardino and San Diego counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves glabrous or sparsely puberulent, especially along margins, blades obovate to elliptic; stems prostrate or decumbent; Colorado.
var. procumbens
1. Leaves glabrate or scabrous to pubescent, blades linear, oblanceolate, or obovate; stems decumbent, ascending, or erect; Colorado, New Mexico, Utah.
→ 2
2. Leaf blades obovate or oblanceolate; stems decumbent to ascending.
→ 3
3. Leaves glabrate or scabrous, blades oblanceolate; Colorado, Utah.
var. crandallii
3. Leaves ± scabrous, blades obovate or oblanceolate; Utah.
var. atratus
2. Leaf blades linear to oblanceolate; stems ascending to erect.
→ 4
4. Stems retrorsely hairy or puberulent, ascending to erect; Colorado, New Mexico.
var. glabrescens
4. Stems puberulent to pubescent, erect; Colorado.
var. ramaleyi
1. Thyrse axes glabrous; stems glabrous, rarely retrorsely hairy; calyx lobes obovate to oblanceolate, rarely lanceolate.
var. heterophyllus
1. Thyrse axes retrorsely hairy; stems retrorsely hairy; calyx lobes ovate to lanceolate.
→ 2
2. Cauline leaves 15–45 × 0.5–4 mm, axillary fascicles present, sometimes absent.
var. australis
2. Cauline leaves (25–)30–100 × 2–12 mm, axillary fascicles absent, rarely present.
var. purdyi
Source FNA vol. 17, p. 101. FNA vol. 17, p. 235.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. crandallii var. atratus, P. crandallii var. crandallii, P. crandallii var. glabrescens, P. crandallii var. procumbens, P. crandallii var. ramaleyi
P. heterophyllus var. australis, P. heterophyllus var. heterophyllus, P. heterophyllus var. purdyi
Name authority A. Nelson: Bull. Torrey Bot. Club 26: 354. (1899) — (as Pentstemon) Lindley: Edwards’s Bot. Reg. 22: plate 1899. (1836) — (as Pentstemon heterophyllum)
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