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Sunset Crater beardtongue, Sunset Crater penstemon

Susanville beardtongue, volcanic beardtongue

Habit Herbs. Subshrubs.
Stems

ascending to erect, 30–60 cm, glaucous.

ascending to erect, 18–70 cm, glandular-pubescent, sometimes densely so distally, not glaucous.

Leaves

basal and proximal cauline 26–56 × 7–15 mm, blade lanceolate to elliptic, base tapered, margins coarsely dentate or serrate, apex obtuse to acute;

cauline 4–7 pairs, short-petiolate or sessile, 30–55 × 5–32 mm, blade ovate to lanceolate, base tapered or auriculate-clasping to connate-perfoliate on distal leaves, margins coarsely dentate, apex acute to acuminate.

cauline, opposite or subopposite, not leathery, glandular-pubescent;

cauline (4–)7–10 pairs, sessile or proximals short-petiolate, 14–60 × 8–20(–30) mm, blade obovate to ovate or lanceolate, base tapered to clasping, margins dentate, dentate-serrate, or serrate, apex acute to acuminate.

Thyrses

interrupted, secund, 8–30 cm, axis glabrous or sparsely glandular-pubescent distally, verticillasters 3–10, cymes 1–3-flowered;

proximal bracts ovate to triangular, 12–35 × 8–32 mm;

peduncles and pedicels ascending to erect, glandular-pubescent.

interrupted or continuous, cylindric, 13–35 cm, axis densely glandular-pubescent, verticillasters 9–18, cymes 1–5-flowered, 2 per node;

proximal bracts ovate to lanceolate, (7–)15–40 × (2–)8–18 mm, margins dentate to dentate-serrate;

peduncles and pedicels ascending to erect, densely glandular-pubescent.

Flowers

calyx lobes ovate to elliptic, 3.5–5.5 × 2.5–3.8 mm, glandular-pubescent;

corolla pink to rose pink, with reddish purple nectar guides, strongly bilabiate, ventricose, 23–27 mm, glandular-pubescent externally, glandular-pubescent internally, also usually moderately white-lanate abaxially, tube 6–10 mm, length 1.6–2 times calyx lobes, throat abruptly inflated, ± constricted at orifice, 9–12 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen sacs explanate, 1.3–1.6 mm, sutures smooth;

staminode 16–18 mm, exserted, 0.4–0.6(–1) mm diam., tip straight, glabrous or distal 2–4 mm moderately lanate, hairs yellow, to 1 mm;

style 17–22 mm, glabrous.

calyx lobes ovate to lanceolate, 3–6 × 0.8–1.6 mm, densely glandular-pubescent;

corolla white to light lavender or light pink, with reddish violet nectar guides, funnelform, 9–11 mm, glandular-pubescent externally, glandular-pubescent internally, tube 3–4 mm, throat slightly inflated, 3–4 mm diam., rounded abaxially;

stamens included, pollen sacs opposite, navicular to subexplanate, 0.5–0.7 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth;

staminode 9–10 mm, included or exserted, 0.2–0.3 mm diam., tip straight, glabrous;

style 7–8 mm.

Capsules

9–12 × 6–7 mm, glabrous.

4–5 × 2.5–3 mm, glabrous.

2n

= 16.

Penstemon clutei

Penstemon sudans

Phenology Flowering Apr–Aug. Flowering Jun–Jul.
Habitat Volcanic cinder fields, pine forests. Igneous soils and crevices, basalt in sagebrush shrublands and pine woodlands.
Elevation 1900–2600 m. (6200–8500 ft.) 1200–1800 m. (3900–5900 ft.)
Distribution
from FNA
AZ
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
[BONAP county map]
Discussion

Penstemon clutei occurs only near Indian Flat and Sunset Crater in south-central Coconino County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon sudans is known from northeastern California (Lassen, Modoc, and Placer counties); it has been reported for Washoe County, Nevada. D. D. Keck (1940b) observed that some plants of P. deustus from southeastern Oregon (Curry, Jackson, and Josephine counties) approach P. sudans in glandularity, though he considered these to be one extreme of the morphologic spectrum exhibited by P. deustus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 249. FNA vol. 17, p. 223.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Synonyms P. pseudospectabilis subsp. clutei P. deustus subsp. sudans
Name authority A. Nelson: Amer. Bot. (Binghamton) 33: 109. (1927) M. E. Jones: Contr. W. Bot. 8: 37. (1898) — (as Pentstemon)
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