Cleveland's beardtongue
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Guadalupe beardtongue
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Herbs. |
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ascending to erect, 30–70 cm, glaucous. |
ascending to erect, 20–50 cm, retrorsely hairy. |
basal and proximal cauline 15–90 × 8–35 mm, blade ovate, base truncate to tapered, margins entire or coarsely dentate, apex acute; cauline 4–7 pairs, short-petiolate or sessile, 10–70 × 8–32 mm, blade cordate to triangular-lanceolate, base tapered or truncate on proximal leaves to connate-perfoliate on distal leaves, margins coarsely dentate, sometimes entire, apex acute. |
basal and cauline, not leathery, glabrous or glabrate, rarely basal retrorsely hairy along midvein; basal and proximal cauline petiolate, 25–95 × 1–6 mm, blade linear, rarely lanceolate, base tapered, margins entire, apex acute; cauline 6–11, sessile or proximals short-petiolate, 22–65(–95) × 1–10(–18) mm, blade lanceolate to linear, base tapered to clasping, margins entire or obscurely serrate, apex acute. |
interrupted, secund, 10–40(–65) cm, axis glabrous or glandular-pubescent, verticillasters 6–12(–22), cymes 2–8-flowered; proximal bracts depressed-ovate to ovate or triangular, 6–45 × 11–26 mm; peduncles and pedicels spreading to ascending or erect, glabrous or glandular-pubescent. |
continuous or ± interrupted, cylindric, 3–16(–26) cm, axis glandular-pubescent, verticillasters 3–8, cymes 2–4-flowered, 2 per node; proximal bracts lanceolate, 15–48 × 3–8 mm; peduncles and pedicels glandular-pubescent. |
calyx lobes ovate, 3.2–6 × 1.6–4 mm, glabrous or glandular-pubescent; corolla red to reddish purple, with or without reddish or reddish purple nectar guides, bilabiate, tubular-funnelform to slightly ventricose, 17–24 mm, glandular-pubescent externally, glabrous internally or glandular-pubescent, sometimes also pilose abaxially, tube 6–9 mm, length 1.8–2 times calyx lobes, throat gradually inflated, slightly constricted or not at orifice, 5–8 mm diam., rounded abaxially; stamens included or longer pair reaching orifice, filaments glabrous, pollen sacs navicular or explanate, 1.2–1.7 mm, sutures smooth or denticulate, teeth to 0.1 mm; staminode 6–11 mm, included, 0.3–0.5 mm diam., tip straight to recurved, glabrous or distal 1–6 mm pilose, hairs yellow, to 0.8 mm, proximal 1–2 mm sometimes glandular-puberulent; style 11–14 mm, glabrous. |
calyx lobes lanceolate, 5–6.5 × 1.2–2 mm, glandular-pubescent; corolla white, sometimes tinged pink or lavender, with reddish violet nectar guides, funnelform, 13–20 mm, glandular-pubescent and sparsely white-villous internally abaxially, tube 4–6 mm, throat gradually inflated, not constricted at orifice, 6–7 mm diam., rounded abaxially; stamens included, pollen sacs opposite, explanate, 0.8–1 mm, dehiscing completely, sutures smooth; staminode 10–14 mm, reaching orifice or exserted, 0.5–0.7 mm diam., tip straight to recurved, distal 3–7 mm sparsely pilose, hairs yellow, to 0.9 mm; style 9–12 mm. |
6–9 × 5–7 mm, glabrous. |
7–9 × 4.5–6 mm. |
= 16. |
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Flowering Mar–Jun. |
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Sandy or gravelly, calcareous soils, prairies. |
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400–800 m. (1300–2600 ft.) |
CA; nw Mexico
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TX |
Varieties 3 (3 in the flora). Penstemon clevelandii is an ancient, stable, diploid hybrid between P. centranthifolius and P. spectabilis (A. D. Wolfe et al. 1998). Penstemon ×parishii A. Gray is a naturally occurring hybrid, also with P. centranthifolius and P. spectabilis as parents; it has been reported from Los Angeles, Riverside, San Bernardino, and San Diego counties, California (D. D. Keck 1937b; Paul Wilson and M. Valenzuela 2002). Keck reported that P. clevelandii var. clevelandii could be distinguished from P. ×parishii by the former’s shorter and narrower corollas, more glandular-pubescent inflorescences, and explanate pollen sacs. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon guadalupensis is known from the southern Cross Timbers and Prairies, Edwards Plateau, and southern Rolling Plains physiographic provinces of central Texas. Pennell distinguished subsp. ernestii by its shorter corollas and broader leaves, but that distinction seems trivial. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Distal cauline leaves sessile, bases connate-perfoliate; pollen sacs navicular, sutures denticulate; peduncles, pedicels, and calyx lobes glabrous. | var. connatus |
1. Distal cauline leaves short-petiolate or sessile, bases tapered or truncate; pollen sacs explanate, sutures smooth; peduncles, pedicels, and calyx lobes glandular-pubescent, rarely glabrous. | → 2 |
2. Corollas glandular-pubescent internally, not pilose or sparsely white- or yellowish pilose internally abaxially; staminodes 9–11 mm. | var. clevelandii |
2. Corollas not glandular-pubescent internally, densely white- or yellowish pilose internally abaxially; staminodes 6–8 mm. | var. mohavensis |
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FNA vol. 17, p. 248. |
FNA vol. 17, p. 137. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. guadalupensis subsp. ernestii |
A. Gray: Proc. Amer. Acad. Arts 11: 94. (1876) — (as Pentstemon clevelandi) |
A. Heller: Contr. Herb. Franklin Marshall Coll. 1: 92, plate 7. (1895) — (as Pentstemon) |
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