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ash beardtongue, ash penstemon, small flower penstemon

Cleveland's beardtongue

Habit Herbs. Herbs.
Stems

ascending, (8–)10–40 cm, glabrous or retrorsely hairy, not glaucous.

ascending to erect, 30–70 cm, glaucous.

Leaves

essentially cauline, basal usually poorly developed, not leathery, glabrous;

basal and proximal cauline 10–40(–65) × 1–6 mm, blade oblanceolate, base tapered, margins entire, apex obtuse to acute;

cauline 3–5 pairs, sessile or proximals short-petiolate, 12–60 × 2–5(–7) mm, blade oblanceolate to lanceolate or linear, base tapered, margins entire, apex acute.

basal and proximal cauline 15–90 × 8–35 mm, blade ovate, base truncate to tapered, margins entire or coarsely dentate, apex acute;

cauline 4–7 pairs, short-petiolate or sessile, 10–70 × 8–32 mm, blade cordate to triangular-lanceolate, base tapered or truncate on proximal leaves to connate-perfoliate on distal leaves, margins coarsely dentate, sometimes entire, apex acute.

Thyrses

interrupted, cylindric, 1–14 cm, axis glabrous or ± puberulent at axils, verticillasters (1 or)2–6, cymes (1–)3–7-flowered, 2 per node;

proximal bracts lanceolate to linear, 8–30(–45) × 1–7 mm, margins entire;

peduncles and pedicels erect, glabrous.

interrupted, secund, 10–40(–65) cm, axis glabrous or glandular-pubescent, verticillasters 6–12(–22), cymes 2–8-flowered;

proximal bracts depressed-ovate to ovate or triangular, 6–45 × 11–26 mm;

peduncles and pedicels spreading to ascending or erect, glabrous or glandular-pubescent.

Flowers

calyx lobes obovate to ovate, 1–1.8 × 0.7–1.1 mm, apex truncate to cuspidate, glabrous;

corolla violet to blue or purple, without nectar guides, funnelform, 7–9(–11) mm, glabrous externally, moderately yellowish or white-pilose internally abaxially, tube 3–4 mm, throat slightly inflated, 2–3 mm diam., 2-ridged abaxially;

stamens included or longer pair reaching orifice, pollen sacs opposite, explanate, 0.3–0.4(–0.5) mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth;

staminode 4–6 mm, included, 0.3–0.4 mm diam., tip straight, distal 0.5–1 mm sparsely to moderately pilose, hairs yellow, to 0.4 mm;

style 5–6 mm.

calyx lobes ovate, 3.2–6 × 1.6–4 mm, glabrous or glandular-pubescent;

corolla red to reddish purple, with or without reddish or reddish purple nectar guides, bilabiate, tubular-funnelform to slightly ventricose, 17–24 mm, glandular-pubescent externally, glabrous internally or glandular-pubescent, sometimes also pilose abaxially, tube 6–9 mm, length 1.8–2 times calyx lobes, throat gradually inflated, slightly constricted or not at orifice, 5–8 mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, filaments glabrous, pollen sacs navicular or explanate, 1.2–1.7 mm, sutures smooth or denticulate, teeth to 0.1 mm;

staminode 6–11 mm, included, 0.3–0.5 mm diam., tip straight to recurved, glabrous or distal 1–6 mm pilose, hairs yellow, to 0.8 mm, proximal 1–2 mm sometimes glandular-puberulent;

style 11–14 mm, glabrous.

Capsules

2.5–4 × 1.8–2.5 mm, glabrous.

6–9 × 5–7 mm, glabrous.

2n

= 16, 32.

= 16.

Penstemon cinicola

Penstemon clevelandii

Phenology Flowering Jun–Aug.
Habitat Dry, volcanic soils in sagebrush openings in pine forests.
Elevation 1000–2600 m. (3300–8500 ft.)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon cinicola occurs along the eastern flank of the Cascade Range in central Oregon (Crook, Deschutes, Douglas, Klamath, and Lake counties) south to northern California (Lassen, Modoc, Shasta, Siskiyou, and Tehama counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Penstemon clevelandii is an ancient, stable, diploid hybrid between P. centranthifolius and P. spectabilis (A. D. Wolfe et al. 1998). Penstemon ×parishii A. Gray is a naturally occurring hybrid, also with P. centranthifolius and P. spectabilis as parents; it has been reported from Los Angeles, Riverside, San Bernardino, and San Diego counties, California (D. D. Keck 1937b; Paul Wilson and M. Valenzuela 2002). Keck reported that P. clevelandii var. clevelandii could be distinguished from P. ×parishii by the former’s shorter and narrower corollas, more glandular-pubescent inflorescences, and explanate pollen sacs.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Distal cauline leaves sessile, bases connate-perfoliate; pollen sacs navicular, sutures denticulate; peduncles, pedicels, and calyx lobes glabrous.
var. connatus
1. Distal cauline leaves short-petiolate or sessile, bases tapered or truncate; pollen sacs explanate, sutures smooth; peduncles, pedicels, and calyx lobes glandular-pubescent, rarely glabrous.
→ 2
2. Corollas glandular-pubescent internally, not pilose or sparsely white- or yellowish pilose internally abaxially; staminodes 9–11 mm.
var. clevelandii
2. Corollas not glandular-pubescent internally, densely white- or yellowish pilose internally abaxially; staminodes 6–8 mm.
var. mohavensis
Source FNA vol. 17, p. 198. FNA vol. 17, p. 248.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. clevelandii var. clevelandii, P. clevelandii var. connatus, P. clevelandii var. mohavensis
Name authority D. D. Keck: Publ. Carnegie Inst. Wash. 520: 294. (1940) A. Gray: Proc. Amer. Acad. Arts 11: 94. (1876) — (as Pentstemon clevelandi)
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