scarlet bugler
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upland beardtongue
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ascending to erect, 30–120 cm, glaucous. |
erect, (10–)30–80 cm, glabrous, not glaucous. |
glabrous, glaucous; cauline 5–11 pairs, short-petiolate or sessile, 40–100 × 10–40 mm, blade ovate to lanceolate, base tapered to auriculate-clasping, margins entire, apex rounded to acute. |
basal and cauline, not leathery, glabrous, not glaucous; basal and proximal cauline 26–140 × 3–19 mm, blade oblanceolate, base tapered, margins entire, apex obtuse to acute; cauline 2–5 pairs, sessile, 50–90 × (3–)6–9(–18) mm, blade oblanceolate to lanceolate, base clasping, apex acute. |
interrupted, secund to ± cylindric, 15–60(–100) cm, axis glabrous or obscurely scabrous, verticillasters 8–18(–22), cymes 1–5(–11)-flowered; proximal bracts ovate to lanceolate, (3–)7–25(–87) × (1–)3–14(–30) mm; peduncles and pedicels erect, glabrous or obscurely scabrous. |
continuous to ± interrupted, secund or cylindric, 5–40 cm, axis glabrous or ± glandular-pubescent, verticillasters (4 or)5–9, cymes 1–5-flowered, 2 per node; proximal bracts lanceolate, 15–54 × 2–15 mm; peduncles and pedicels glabrous or glandular-pubescent, peduncles to 11 mm, pedicels 1–10 mm. |
calyx lobes ovate to lanceolate, 3–6 × 2–3 mm, margins erose, glabrous; corolla scarlet, without nectar guides, nearly radially symmetric, weakly bilabiate, salverform, 25–33 mm, glabrous externally, glabrous internally, tube 8–10 mm, throat slightly inflated, 4.5–6 mm diam., rounded abaxially; stamens: longer pair reaching orifice, pollen sacs explanate, 0.9–1.2 mm, sutures smooth; staminode 13–14 mm, terete to slightly flattened distally, 0.1–0.3 mm diam., tip straight, glabrous; style 18–27 mm, exserted. |
calyx lobes ovate, 3.5–8 × 1.8–3 mm, margins erose, glabrous or glandular-pubescent; corolla blue to bluish violet, with or without reddish purple nectar guides, ventricose, 17–25(–30) mm, glabrous or glandular-pubescent externally, glabrous or ± glandular-pubescent internally abaxially, tube 5–8 mm, throat gradually inflated, slightly constricted at orifice, 6–8 mm diam., rounded abaxially; stamens: longer pair reaching orifice or slightly exserted, pollen sacs opposite, navicular, 1.2–1.8 mm, dehiscing incompletely, connective not splitting, sides sparsely to moderately hispid or pubescent, hairs whitish, to 0.3 mm, sutures denticulate, teeth to 0.1 mm; staminode 12–15 mm, reaching orifice or slightly exserted, 0.4–0.6 mm diam., tip straight to recurved, distal 3–5 mm sparsely villous, hairs yellow, to 1 mm; style 15–18 mm. |
8–11 × 6–7 mm. |
8–12 × 4–6 mm. |
= 16. |
= 16. |
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Flowering Feb–Jul. |
Flowering Jun–Aug. |
Hillsides, chaparral, oak, pinyon-juniper, and Joshua tree woodlands, coastal sage scrub, pine forests. |
Sagebrush shrublands, openings in pine forests. |
0–1800 m. (0–5900 ft.) |
2400–3600 m. (7900–11800 ft.) |
CA; Mexico (Baja California)
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CO; WY |
Penstemon ×parishii A. Gray, a naturally occurring hybrid between P. centranthifolius and P. spectabilis (A. D. Wolfe and W. J. Elisens 1993), has been reported from Los Angeles, Riverside, San Bernardino, and San Diego counties, California (D. D. Keck 1937; Paul Wilson and M. Valenzuela 2002). A wild hybrid between P. centranthifolius and P. eatonii also was reported from San Bernardino County, California (Wilson and Valenzuela). Penstemon ×dubius Davidson was described from Mount Lowe in the San Gabriel Mountains, Los Angeles County, California, growing with P. centranthifolius and P. grinnellii. Hybridization between P. centranthifolius and P. grinnellii has been documented using allozyme and DNA data (Wolfe and Elisens 1993, 1994). Three southern and central Californian tribes of Native Americans used Penstemon centranthifolius for drugs, food, and decorations (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon saxosorum is known in north-central and northwestern Colorado and south-central Wyoming. Populations in the Bridger Basin in Moffat County, Colorado, and Daggett and Uintah counties, Utah, lie between the main ranges of P. saxosorum and P. subglaber. Plants in those populations have pollen sacs 0.7–0.9 mm; corollas sparsely glandular and 18–20 mm; and corolla throats 5–7 mm in diameter. M. L. Moorman (1982) believed that they represented an undescribed species. Pending further study, Colorado plants are referred to P. saxosorum and Utah plants to P. subglaber. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 151. |
FNA vol. 17, p. 178. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
Chelone centranthifolia |
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(Bentham) Bentham: Scroph. Ind., 7. (1835) — (as Pentstemon centranthifolium) |
Pennell: Contr. U.S. Natl. Herb. 20: 349. (1920) |
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