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scarlet bugler

Berry's penstemon, mountain pride, Newberry's beardtongue

Habit Subshrubs, cespitose.
Stems

ascending to erect, 30–120 cm, glaucous.

ascending to erect, (5–)10–32 cm, glabrous or retrorsely hairy, not glaucous.

Leaves

glabrous, glaucous;

cauline 5–11 pairs, short-petiolate or sessile, 40–100 × 10–40 mm, blade ovate to lanceolate, base tapered to auriculate-clasping, margins entire, apex rounded to acute.

persistent, 4–9 pairs, distals distinctly to slightly smaller than proximals, short-petiolate or sessile, (5–)9–42 × 4–14(–18) mm, blade elliptic to obovate, base tapered, cuneate, or cordate-clasping, margins ± serrulate or serrulate-denticulate to serrate, apex rounded to obtuse or acute, glabrous, glaucous.

Thyrses

interrupted, secund to ± cylindric, 15–60(–100) cm, axis glabrous or obscurely scabrous, verticillasters 8–18(–22), cymes 1–5(–11)-flowered;

proximal bracts ovate to lanceolate, (3–)7–25(–87) × (1–)3–14(–30) mm;

peduncles and pedicels erect, glabrous or obscurely scabrous.

continuous, ± secund, 2–20 cm, axis glandular-pubescent, verticillasters 4–12, cymes 1-flowered;

proximal bracts ovate to lanceolate, 4–11(–20) × 2–8(–15) mm;

peduncles and pedicels ascending, glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, 3–6 × 2–3 mm, margins erose, glabrous;

corolla scarlet, without nectar guides, nearly radially symmetric, weakly bilabiate, salverform, 25–33 mm, glabrous externally, glabrous internally, tube 8–10 mm, throat slightly inflated, 4.5–6 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, pollen sacs explanate, 0.9–1.2 mm, sutures smooth;

staminode 13–14 mm, terete to slightly flattened distally, 0.1–0.3 mm diam., tip straight, glabrous;

style 18–27 mm, exserted.

calyx lobes lanceolate, 6–10 × 2–2.5 mm, glandular-pubescent;

corolla pink, red, scarlet, lavender, or purple, essentially unlined internally but abaxial ridges sometimes pink to whitish, not personate, funnelform, 22–35 mm, glabrous externally, moderately to densely white-lanate internally abaxially, tube 6–9 mm, throat 5–12 mm diam.;

stamens included or longer pair exserted, pollen sacs 0.8–1.7 mm;

staminode 8–14 mm, slightly flattened distally, 0.1–0.3 mm diam., tip straight, distal 2–8 mm pubescent to lanate, hairs white or yellowish, to 1 mm;

style 20–24 mm.

Capsules

8–11 × 6–7 mm.

(8–)10–13 × (4–)5–8 mm.

2n

= 16.

Penstemon centranthifolius

Penstemon newberryi

Phenology Flowering Feb–Jul.
Habitat Hillsides, chaparral, oak, pinyon-juniper, and Joshua tree woodlands, coastal sage scrub, pine forests.
Elevation 0–1800 m. (0–5900 ft.)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon ×parishii A. Gray, a naturally occurring hybrid between P. centranthifolius and P. spectabilis (A. D. Wolfe and W. J. Elisens 1993), has been reported from Los Angeles, Riverside, San Bernardino, and San Diego counties, California (D. D. Keck 1937; Paul Wilson and M. Valenzuela 2002). A wild hybrid between P. centranthifolius and P. eatonii also was reported from San Bernardino County, California (Wilson and Valenzuela). Penstemon ×dubius Davidson was described from Mount Lowe in the San Gabriel Mountains, Los Angeles County, California, growing with P. centranthifolius and P. grinnellii. Hybridization between P. centranthifolius and P. grinnellii has been documented using allozyme and DNA data (Wolfe and Elisens 1993, 1994).

Three southern and central Californian tribes of Native Americans used Penstemon centranthifolius for drugs, food, and decorations (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Penstemon newberryi is known from the Sierra Nevada and Cascade ranges of California, Nevada, and Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas pink to lavender or purple, 27–35 mm; stamens included, pollen sacs 1.2–1.7 mm.
var. berryi
1. Corollas red, scarlet, or purple, 22–30 mm; stamens: longer pair exserted, pollen sacs 0.8–1.3 mm.
→ 2
2. Corollas red to scarlet; distal leaves distinctly smaller than proximals.
var. newberryi
2. Corollas purple; distal leaves slightly smaller than proximals.
var. sonomensis
Source FNA vol. 17, p. 151. FNA vol. 17, p. 90.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides Plantaginaceae > Penstemon > subg. Dasanthera > sect. Erianthera
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. newberryi var. berryi, P. newberryi var. newberryi, P. newberryi var. sonomensis
Synonyms Chelone centranthifolia
Name authority (Bentham) Bentham: Scroph. Ind., 7. (1835) — (as Pentstemon centranthifolium) A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 6(3): 82, plate 14. (1858) — (as Pentstemon)
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