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scarlet bugler

Arkansas beardtongue

Habit Herbs.
Stems

ascending to erect, 30–120 cm, glaucous.

ascending to erect, (18–)25–45(–61) cm, retrorsely hairy, sometimes also ± glandular-pubescent distally, not glaucous.

Leaves

glabrous, glaucous;

cauline 5–11 pairs, short-petiolate or sessile, 40–100 × 10–40 mm, blade ovate to lanceolate, base tapered to auriculate-clasping, margins entire, apex rounded to acute.

basal and cauline, basal often withering by anthesis, not leathery, glabrous or sparsely retrorsely hairy, especially along midvein and margins, abaxially, glabrous or sparsely puberulent adaxially;

basal and proximal cauline 20–95 × 5–28 mm, blade spatulate or obovate to ovate, base tapered, margins subentire to ± serrate or dentate, apex rounded to obtuse or acute;

cauline (6–)8–12 pairs, short-petiolate or sessile, 22–80 × 4–20 mm, blade lanceolate, base tapered to clasping, margins ± serrate or dentate, apex acute to acuminate.

Thyrses

interrupted, secund to ± cylindric, 15–60(–100) cm, axis glabrous or obscurely scabrous, verticillasters 8–18(–22), cymes 1–5(–11)-flowered;

proximal bracts ovate to lanceolate, (3–)7–25(–87) × (1–)3–14(–30) mm;

peduncles and pedicels erect, glabrous or obscurely scabrous.

interrupted, conic, 6–24 cm, axis retrorsely hairy and ± glandular-pubescent, verticillasters 4–8, cymes 3–13-flowered, 2 per node;

proximal bracts lanceolate, 6–40 × 2–12 mm, margins entire or ± dentate;

peduncles and pedicels ascending, retrorsely hairy and ± glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, 3–6 × 2–3 mm, margins erose, glabrous;

corolla scarlet, without nectar guides, nearly radially symmetric, weakly bilabiate, salverform, 25–33 mm, glabrous externally, glabrous internally, tube 8–10 mm, throat slightly inflated, 4.5–6 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, pollen sacs explanate, 0.9–1.2 mm, sutures smooth;

staminode 13–14 mm, terete to slightly flattened distally, 0.1–0.3 mm diam., tip straight, glabrous;

style 18–27 mm, exserted.

calyx lobes ovate, 2.8–4.5 × 1.1–2 mm, sparsely glandular-pubescent;

corolla white, with reddish purple nectar guides, tubular to tubular-funnelform, 15–19 mm, glandular-pubescent externally, moderately whitish lanate internally abaxially, tube 4–6 mm, throat slightly inflated, 4–7 mm diam., 2-ridged abaxially;

stamens included, pollen sacs opposite, navicular, 0.8–1 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate;

staminode 10–12 mm, exserted, 0.2–0.4 mm diam., tip straight to slightly recurved, distal 5–7 mm ± villous, hairs yellow or golden yellow, to 1.2 mm;

style 9–11 mm.

Capsules

8–11 × 6–7 mm.

4–7 × 4–5 mm, glabrous.

2n

= 16.

= 16.

Penstemon centranthifolius

Penstemon arkansanus

Phenology Flowering Feb–Jul. Flowering Apr–Jun.
Habitat Hillsides, chaparral, oak, pinyon-juniper, and Joshua tree woodlands, coastal sage scrub, pine forests. Rocky oak-hickory woodlands, glades, rocky stream bottoms.
Elevation 0–1800 m. (0–5900 ft.) 60–300 m. (200–1000 ft.)
Distribution
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; MO; OK
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon ×parishii A. Gray, a naturally occurring hybrid between P. centranthifolius and P. spectabilis (A. D. Wolfe and W. J. Elisens 1993), has been reported from Los Angeles, Riverside, San Bernardino, and San Diego counties, California (D. D. Keck 1937; Paul Wilson and M. Valenzuela 2002). A wild hybrid between P. centranthifolius and P. eatonii also was reported from San Bernardino County, California (Wilson and Valenzuela). Penstemon ×dubius Davidson was described from Mount Lowe in the San Gabriel Mountains, Los Angeles County, California, growing with P. centranthifolius and P. grinnellii. Hybridization between P. centranthifolius and P. grinnellii has been documented using allozyme and DNA data (Wolfe and Elisens 1993, 1994).

Three southern and central Californian tribes of Native Americans used Penstemon centranthifolius for drugs, food, and decorations (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon arkansanus is centered in the Interior Highlands. The species is most similar to P. pallidus, from which it differs by its narrower, less hairy, and more numerous cauline leaves. Some specimens from the narrow zone of sympatry in north-central Arkansas and south-central Missouri exhibit varying degrees of morphologic intermediacy. Penstemon arkansanus has been reported from Illinois (R. H. Mohlenbrock 1986) and Texas (S. D. Jones et al. 1997); those reports seem unlikely.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 151. FNA vol. 17, p. 193.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Synonyms Chelone centranthifolia P. multicaulis, P. pallidus subsp. arkansanus, P. wherryi
Name authority (Bentham) Bentham: Scroph. Ind., 7. (1835) — (as Pentstemon centranthifolium) Pennell: Proc. Acad. Nat. Sci. Philadelphia 73: 493. (1922)
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