Penstemon cardwellii |
Penstemon subg. Dasanthera |
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Cardwell's beardtongue, Cardwell's penstemon |
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Habit | Shrubs, sometimes cespitose. | |
Stems | ascending to erect, 10–27 cm, retrorsely hairy or puberulent, not glaucous. |
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Leaves | persistent, 6–9 pairs, distals usually distinctly smaller than proximals, short-petiolate or sessile, 15–55 × 4–16 mm, blade elliptic, base tapered, margins subentire or serrate, apex rounded to obtuse or acute, glabrous, ± glaucous. |
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Thyrses | ± interrupted, secund, 5–14 cm, axis retrorsely hairy proximally, retrorsely hairy and glandular-pubescent distally, verticillasters 3–7, cymes 1-flowered; proximal bracts ovate to lanceolate, 6–18 × 3–12 mm, margins serrate, sometimes entire; peduncles and pedicels ascending to erect, glandular-pubescent, sometimes also puberulent. |
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Flowers | calyx lobes ovate to lanceolate, 8–14 × 2–3.1 mm, glandular-pubescent; corolla purple to violet, essentially unlined internally but abaxial ridges usually light purple or white, not personate, funnelform, (25–)30–38 mm, glabrous externally, sparsely white-lanate internally abaxially, tube 6–8 mm, throat 9–10 mm diam.; stamens included, pollen sacs 1.5–1.7 mm; staminode 11–13 mm, essentially terete distally, 0.1 mm diam., tip straight, distal 4–6 mm moderately to densely villous, hairs white or yellowish, to 1.5 mm; style 25–29 mm. |
pollen sacs usually explanate, dehiscing completely, connective splitting, sides usually densely lanate, (subexplanate, sides glabrous in P. personatus), sutures smooth; staminode included, length usually 1/3–3/4 times corolla throat. |
Capsules | 8–11 × 4–6 mm. |
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Seeds | angled, wings absent or narrow. |
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2n | = 16. |
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Penstemon cardwellii |
Penstemon subg. Dasanthera |
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Phenology | Flowering May–Aug. | |
Habitat | Rock outcrops, pumice and talus slopes, forest clearings, roadcuts. | |
Elevation | 400–1900 m. (1300–6200 ft.) | |
Distribution |
OR; WA
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w North America |
Discussion | Penstemon cardwellii occurs primarily on the west slope of the Cascade Range from southwestern Oregon to northwestern Washington, with isolated populations east of the crest. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 10 (10 in the flora). Subgenus Dasanthera appears to be the basal lineage in the clade corresponding to Penstemon (A. D. Wolfe et al. 2006). As circumscribed here, subg. Dasanthera comprises a single section, sect. Erianthera. Morphologic and molecular data indicate that the persistent-leaved lineage of the Sierra-Cascade is derived from the mostly deciduous-leaved lineage of the Rocky Mountains (A. D. Every 1977; S. L. Datwyler and Wolfe 2004). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 86. | FNA vol. 17, p. 85. |
Parent taxa | Plantaginaceae > Penstemon > subg. Dasanthera > sect. Erianthera | Plantaginaceae > Penstemon |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Dasanthera | |
Name authority | Howell: Fl. N.W. Amer., 510. (1901) — (as Pentstemon) | (Rafinesque) Pennell: Contr. U.S. Natl. Herb. 20: 327. (1920) |
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