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Cardwell's beardtongue, Cardwell's penstemon

Blue Ridge beardtongue, Small's beardtongue

Habit Shrubs, sometimes cespitose. Herbs.
Stems

ascending to erect, 10–27 cm, retrorsely hairy or puberulent, not glaucous.

ascending to erect, 35–80 cm, puberulent or retrorsely hairy proximally, retrorsely hairy and sparsely glandular-pubescent distally, not glaucous.

Leaves

persistent, 6–9 pairs, distals usually distinctly smaller than proximals, short-petiolate or sessile, 15–55 × 4–16 mm, blade elliptic, base tapered, margins subentire or serrate, apex rounded to obtuse or acute, glabrous, ± glaucous.

basal and cauline, basal sometimes withering by anthesis, not leathery, glabrous or puberulent along midveins and, sometimes, on proximal parts of blade;

basal and proximal cauline 55–170 × 15–60 mm, blade triangular-ovate to cordate or lanceolate, base tapered, margins crenate to sharply serrate, apex obtuse to acute;

cauline 4–6 pairs, sessile, 66–105 × 12–48 mm, blade ovate to lanceolate, proximals sometimes lyrate, base truncate to broadly clasping, margins crenate to sharply serrate, apex acute.

Thyrses

± interrupted, secund, 5–14 cm, axis retrorsely hairy proximally, retrorsely hairy and glandular-pubescent distally, verticillasters 3–7, cymes 1-flowered;

proximal bracts ovate to lanceolate, 6–18 × 3–12 mm, margins serrate, sometimes entire;

peduncles and pedicels ascending to erect, glandular-pubescent, sometimes also puberulent.

interrupted, narrowly conic, (6–)10–28 cm, axis puberulent and glandular-pubescent, verticillasters 4–7, cymes (3–)5–12-flowered, 2 per node;

proximal bracts lanceolate, 56–130 × 23–55 mm, margins sharply serrate;

peduncles and pedicels ascending to erect, puberulent and glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, 8–14 × 2–3.1 mm, glandular-pubescent;

corolla purple to violet, essentially unlined internally but abaxial ridges usually light purple or white, not personate, funnelform, (25–)30–38 mm, glabrous externally, sparsely white-lanate internally abaxially, tube 6–8 mm, throat 9–10 mm diam.;

stamens included, pollen sacs 1.5–1.7 mm;

staminode 11–13 mm, essentially terete distally, 0.1 mm diam., tip straight, distal 4–6 mm moderately to densely villous, hairs white or yellowish, to 1.5 mm;

style 25–29 mm.

calyx lobes ovate to lanceolate, 3.5–5 × 1.2–2.1 mm, glandular-pubescent;

corolla lavender to violet or purple, with violet nectar guides, funnelform to ventricose-ampliate, 28–35 mm, glandular-pubescent externally, moderately white-lanate internally abaxially, tube 7–8 mm, throat abruptly inflated, 9–16 mm diam., 2-ridged abaxially;

stamens included, pollen sacs opposite, navicular, 1.3–1.5 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate;

staminode 15–18 mm, included, 0.4–0.6 mm diam., tip straight, distal 13–15 mm ± pilose, hairs yellow, to 2 mm;

style 17–20 mm.

Capsules

8–11 × 4–6 mm.

7–10 × 4–5 mm, glabrous.

2n

= 16.

= 16.

Penstemon cardwellii

Penstemon smallii

Phenology Flowering May–Aug. Flowering Apr–Jul.
Habitat Rock outcrops, pumice and talus slopes, forest clearings, roadcuts. Rocky slopes, bluffs, cliffs.
Elevation 400–1900 m. (1300–6200 ft.) 200–1200 m. (700–3900 ft.)
Distribution
from FNA
OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; GA; NC; SC; TN
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon cardwellii occurs primarily on the west slope of the Cascade Range from southwestern Oregon to northwestern Washington, with isolated populations east of the crest.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon smallii is known from the southern Appalachians. Foliose inflorescence bracts, truncate or cordate cauline leaf bases, and lavender to purple corollas distinguish it from other eastern penstemons.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 86. FNA vol. 17, p. 222.
Parent taxa Plantaginaceae > Penstemon > subg. Dasanthera > sect. Erianthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority Howell: Fl. N.W. Amer., 510. (1901) — (as Pentstemon) A. Heller: Bull. Torrey Bot. Club 21: 25. (1894) — (as Pentstemon)
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