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Cardwell's beardtongue, Cardwell's penstemon

Palmer's beardtongue, Palmer's penstemon, scented beardtongue, scented penstemon

Habit Shrubs, sometimes cespitose. Herbs.
Stems

ascending to erect, 10–27 cm, retrorsely hairy or puberulent, not glaucous.

ascending to erect, 43–140 cm, glaucous.

Leaves

persistent, 6–9 pairs, distals usually distinctly smaller than proximals, short-petiolate or sessile, 15–55 × 4–16 mm, blade elliptic, base tapered, margins subentire or serrate, apex rounded to obtuse or acute, glabrous, ± glaucous.

basal and proximal cauline (28–)52–120(–145) × (5–)25–50(–60) mm, blade ovate, base tapered, margins coarsely dentate, apex obtuse to acute;

cauline 5–8 pairs, sessile or connate-perfoliate, 25–130 × 7–45 mm, blade ovate to triangular, base of distals auriculate-clasping or connate-perfoliate, margins coarsely dentate, distals sometimes entire, apex acute.

Thyrses

± interrupted, secund, 5–14 cm, axis retrorsely hairy proximally, retrorsely hairy and glandular-pubescent distally, verticillasters 3–7, cymes 1-flowered;

proximal bracts ovate to lanceolate, 6–18 × 3–12 mm, margins serrate, sometimes entire;

peduncles and pedicels ascending to erect, glandular-pubescent, sometimes also puberulent.

interrupted, rarely continuous, secund, 7–63 cm, axis glabrous or glandular-pubescent, verticillasters (4–)7–17, cymes (1 or)2–5-flowered;

proximal bracts ovate to lanceolate, 8–40 × 4–38 mm;

peduncles and pedicels ascending to erect, glabrous or glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, 8–14 × 2–3.1 mm, glandular-pubescent;

corolla purple to violet, essentially unlined internally but abaxial ridges usually light purple or white, not personate, funnelform, (25–)30–38 mm, glabrous externally, sparsely white-lanate internally abaxially, tube 6–8 mm, throat 9–10 mm diam.;

stamens included, pollen sacs 1.5–1.7 mm;

staminode 11–13 mm, essentially terete distally, 0.1 mm diam., tip straight, distal 4–6 mm moderately to densely villous, hairs white or yellowish, to 1.5 mm;

style 25–29 mm.

calyx lobes ovate, (3.4–)4.2–6(–7.5) × 1.9–3.6 mm, glabrous or glandular-pubescent;

corolla pale pink to white or pinkish lavender, with reddish purple nectar guides, strongly bilabiate, ventricose-ampliate, 25–38 mm, glandular-pubescent externally, glandular-pubescent internally, sometimes also sparsely white-lanate abaxially, tube 4–8 mm, length 1–1.4 times calyx lobes, throat abruptly inflated, constricted at orifice, 12–21 mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen sacs navicular, 1.6–2.4 mm, sutures smooth;

staminode 25–30 mm, exserted, 0.9–1.1 mm diam., tip strongly recurved to coiled, distal 8–10 mm, and especially tip, moderately to densely lanate, hairs yellow, to 3.5 mm, proximal 6–8 mm glandular-puberulent;

style 19–23 mm, glabrous or sparsely glandular-pubescent proximally.

Capsules

8–11 × 4–6 mm.

11–16 × 6–8 mm, sparsely glandular-pubescent distally.

2n

= 16.

Penstemon cardwellii

Penstemon palmeri

Phenology Flowering May–Aug.
Habitat Rock outcrops, pumice and talus slopes, forest clearings, roadcuts.
Elevation 400–1900 m. (1300–6200 ft.)
Distribution
from FNA
OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; NM; NV; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon cardwellii occurs primarily on the west slope of the Cascade Range from southwestern Oregon to northwestern Washington, with isolated populations east of the crest.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Penstemon ×bryantiae D. D. Keck, a hybrid between P. palmeri and P. spectabilis, was described based on plants that volunteered in the Rancho Santa Ana Botanic Garden in 1935 (D. D. Keck 1937b). Penstemon ×mirus A. Nelson, a putative hybrid between P. eatonii and P. palmeri, has been reported from Arizona (A. Nelson 1938).

A poultice prepared from Penstemon palmeri is applied to snakebites by the Kayenta Navajo of northeastern Arizona (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corolla tubes 7–8 mm; distal cauline leaves sessile, sometimes connate-perfoliate.
var. macranthus
1. Corolla tubes 4–6 mm; distal cauline leaves connate-perfoliate.
→ 2
2. Peduncles, pedicels, and calyx lobes glandular-pubescent; pollen sacs 1.8–2.4 mm.
var. palmeri
2. Peduncles, pedicels, and calyx lobes glabrous; pollen sacs 1.6–2(–2.2) mm.
var. eglandulosus
Source FNA vol. 17, p. 86. FNA vol. 17, p. 252.
Parent taxa Plantaginaceae > Penstemon > subg. Dasanthera > sect. Erianthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. palmeri var. eglandulosus, P. palmeri var. macranthus, P. palmeri var. palmeri
Name authority Howell: Fl. N.W. Amer., 510. (1901) — (as Pentstemon) A. Gray: Proc. Amer. Acad. Arts 7: 379. (1868) — (as Pentstemon)
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