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Cardwell's beardtongue, Cardwell's penstemon

Penstemon anguineus

Siskiyou beardtongue, tongue leaf penstemon

Habit Shrubs, sometimes cespitose. Herbs.
Stems

ascending to erect, 10–27 cm, retrorsely hairy or puberulent, not glaucous.

ascending to erect, (10–)30–80 cm, glabrous, not glaucous.

Leaves

persistent, 6–9 pairs, distals usually distinctly smaller than proximals, short-petiolate or sessile, 15–55 × 4–16 mm, blade elliptic, base tapered, margins subentire or serrate, apex rounded to obtuse or acute, glabrous, ± glaucous.

basal and cauline, not leathery, glabrous;

basal and proximal cauline 30–150 × 6–40 mm, blade ovate to elliptic or oval, base tapered, margins serrate to dentate, sometimes denticulate or entire, apex obtuse to acute;

cauline 2–4 pairs, sessile or proximals short-petiolate, 11–90 × 5–50 mm, blade oblanceolate to oblong or triangular-ovate, base tapered to cordate-clasping, margins serrate to denticulate, rarely entire, apex obtuse to acute.

Thyrses

± interrupted, secund, 5–14 cm, axis retrorsely hairy proximally, retrorsely hairy and glandular-pubescent distally, verticillasters 3–7, cymes 1-flowered;

proximal bracts ovate to lanceolate, 6–18 × 3–12 mm, margins serrate, sometimes entire;

peduncles and pedicels ascending to erect, glandular-pubescent, sometimes also puberulent.

interrupted, sometimes continuous, cylindric, (4–)8–40 cm, axis glandular-pubescent, verticillasters 3–10, cymes (2 or)3–15-flowered, 2 per node;

proximal bracts ovate to triangular, 10–85 × 7–40 mm, margins serrate or entire;

peduncles and pedicels ascending, glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, 8–14 × 2–3.1 mm, glandular-pubescent;

corolla purple to violet, essentially unlined internally but abaxial ridges usually light purple or white, not personate, funnelform, (25–)30–38 mm, glabrous externally, sparsely white-lanate internally abaxially, tube 6–8 mm, throat 9–10 mm diam.;

stamens included, pollen sacs 1.5–1.7 mm;

staminode 11–13 mm, essentially terete distally, 0.1 mm diam., tip straight, distal 4–6 mm moderately to densely villous, hairs white or yellowish, to 1.5 mm;

style 25–29 mm.

calyx lobes lanceolate, 4–6.5 × 0.9–2 mm, glandular-pubescent;

corolla light blue to lavender, purple, or violet, with violet nectar guides, ampliate, 13–18 mm, glandular-pubescent externally, glabrous or sparsely white-lanate internally abaxially, tube 5–7 mm, throat gradually inflated, 4–6 mm diam., 2-ridged abaxially;

stamens included, pollen sacs opposite, navicular, 0.8–1.2 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate;

staminode 10–13 mm, exserted, 0.2–0.4 mm diam., tip straight to slightly recurved, distal 2–6 mm glabrous or sparsely pilose, hairs yellow, to 0.6 mm;

style 10–13 mm.

Capsules

8–11 × 4–6 mm.

5–7 × 3–4 mm, glabrous.

2n

= 16.

= 16.

Penstemon cardwellii

Penstemon anguineus

Phenology Flowering May–Aug. Flowering Jun–Aug.
Habitat Rock outcrops, pumice and talus slopes, forest clearings, roadcuts. Rocky, clayey, or humus soils, openings in coniferous forests.
Elevation 400–1900 m. (1300–6200 ft.) 600–2000 m. (2000–6600 ft.)
Distribution
from FNA
OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon cardwellii occurs primarily on the west slope of the Cascade Range from southwestern Oregon to northwestern Washington, with isolated populations east of the crest.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon anguineus occurs primarily in the Cascade, Klamath, and North Coast ranges from southwestern Oregon (Curry, Douglas, Jackson, Josephine, and Klamath counties) south into northwestern California (Del Norte, El Dorado, Glenn, Humboldt, Mendocino, Siskiyou, Tehama, and Trinity counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 86. FNA vol. 17, p. 191.
Parent taxa Plantaginaceae > Penstemon > subg. Dasanthera > sect. Erianthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority Howell: Fl. N.W. Amer., 510. (1901) — (as Pentstemon) Eastwood: Bull. Torrey Bot. Club 32: 208. (1905) — (as Pentstemon)
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