Cardwell's beardtongue, Cardwell's penstemon
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Alberta beardtongue, Alberta penstemon
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Shrubs, sometimes cespitose. |
Herbs. |
ascending to erect, 10–27 cm, retrorsely hairy or puberulent, not glaucous. |
ascending to erect, (10–)15–38 cm, glabrous or sparsely retrorsely hairy, sometimes also sparsely glandular-pubescent. |
persistent, 6–9 pairs, distals usually distinctly smaller than proximals, short-petiolate or sessile, 15–55 × 4–16 mm, blade elliptic, base tapered, margins subentire or serrate, apex rounded to obtuse or acute, glabrous, ± glaucous. |
basal and cauline, ± leathery or not, glabrous, sometimes sparsely retrorsely hairy along midvein abaxially, not glaucous; basal and proximal cauline (13–)40–90 × (3–)6–18 mm, blade ovate to oblanceolate or lanceolate, base tapered, margins ± serrate or dentate, apex obtuse to acute; cauline 2–6 pairs, short-petiolate or sessile, 15–55 × 3–10 mm, blade lanceolate, proximals sometimes oblanceolate to oblong, base tapered or truncate, margins entire or ± serrate to ± dentate, apex obtuse to acute. |
± interrupted, secund, 5–14 cm, axis retrorsely hairy proximally, retrorsely hairy and glandular-pubescent distally, verticillasters 3–7, cymes 1-flowered; proximal bracts ovate to lanceolate, 6–18 × 3–12 mm, margins serrate, sometimes entire; peduncles and pedicels ascending to erect, glandular-pubescent, sometimes also puberulent. |
continuous or ± interrupted, cylindric, 4–19 cm, axis glabrous or ± glandular-pubescent, verticillasters 4–8, cymes (1 or)2–6-flowered, 2 per node; proximal bracts lanceolate, 9–40 × 3–15 mm, margins entire or ± serrate to dentate; peduncles and pedicels erect, sometimes ascending, glandular-pubescent. |
calyx lobes ovate to lanceolate, 8–14 × 2–3.1 mm, glandular-pubescent; corolla purple to violet, essentially unlined internally but abaxial ridges usually light purple or white, not personate, funnelform, (25–)30–38 mm, glabrous externally, sparsely white-lanate internally abaxially, tube 6–8 mm, throat 9–10 mm diam.; stamens included, pollen sacs 1.5–1.7 mm; staminode 11–13 mm, essentially terete distally, 0.1 mm diam., tip straight, distal 4–6 mm moderately to densely villous, hairs white or yellowish, to 1.5 mm; style 25–29 mm. |
calyx lobes ovate, 2.8–4.8 × 0.9–1.8 mm, glandular-pubescent; corolla blue to light blue or violet, with reddish purple nectar guides, funnelform, 13–20 mm, glandular-pubescent externally, sparsely white-villous internally abaxially, tube 4–5 mm, throat gradually inflated, 4–5 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, subexplanate to explanate, 0.5–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate; staminode 8–9 mm, included, 0.2–0.3 mm diam., tip straight to recurved, distal 0.5–2 mm ± pilose, hairs golden yellow, to 0.8 mm; style 9–11 mm. |
8–11 × 4–6 mm. |
4–6 × 3–4 mm, glabrous. |
= 16. |
= 16. |
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Flowering May–Aug. |
Flowering May–Aug. |
Rock outcrops, pumice and talus slopes, forest clearings, roadcuts. |
Gravelly slopes, cliffs, rocky clearings, roadcuts in coniferous forests. |
400–1900 m. (1300–6200 ft.) |
800–2800 m. (2600–9200 ft.) |
OR; WA
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ID; MT; AB; BC
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Penstemon cardwellii occurs primarily on the west slope of the Cascade Range from southwestern Oregon to northwestern Washington, with isolated populations east of the crest. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon caelestinus has been treated as a synonym of P. albertinus (D. D. Keck 1945; A. Cronquist 1959). D. V. Clark (1971) grew these and related species in transplant gardens and considered P. caelestinus to be distinct based on flower color, leaf shape and color, phenology, and habitat. A broad zone of hybridization between P. albertinus and P. wilcoxii exists in east-central Idaho and western Montana, especially in habitats disturbed by humans (Clark). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 86. |
FNA vol. 17, p. 190. |
Plantaginaceae > Penstemon > subg. Dasanthera > sect. Erianthera |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. caelestinus |
Howell: Fl. N.W. Amer., 510. (1901) — (as Pentstemon) |
Greene: Leafl. Bot. Observ. Crit. 1: 167. (1906) — (as Pentstemon) |
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