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cardinal beardtongue, scarlet beardtongue

Penstemon rattanii

Rattan's beardtongue, Rattan's penstemon

Habit Herbs.
Stems

ascending to erect, 32–100 cm, glabrous, slightly glaucous.

ascending to erect, (15–)30–120 cm, glabrous, sometimes glabrous proximally, glandular-pubescent distally, not glaucous.

Leaves

glabrous, not glaucous;

basal and proximal cauline 40–80(–150) × 14–40(–60) mm, blade spatulate to obovate, oblanceolate, or elliptic, base tapered, margins entire, apex rounded to obtuse;

cauline 3–7 pairs, sessile, (11–)20–135 × 9–50 mm, blade spatulate to elliptic, ovate, or subcordate, base truncate to broadly cordate-clasping, apex rounded to acute or mucronate.

basal and cauline, not leathery, glabrous;

basal and proximal cauline 30–250 × 7–50 mm, blade elliptic to lanceolate, base tapered, margins serrate to dentate, apex obtuse to acute;

cauline 2–5 pairs, short-petiolate or sessile, (10–)20–75(–110) × 5–30(–38) mm, blade oblong to triangular-ovate, base truncate to cordate-clasping distally, margins serrate to dentate, apex obtuse to acute.

Thyrses

secund, (3–)6–44 cm, axis glabrous, verticillasters 3–15, cymes 2–4(–8)-flowered;

proximal bracts suborbiculate, ovate, or lanceolate, (4–)16–60 × (2–)5–45 mm;

peduncles and pedicels erect, glabrous.

interrupted, cylindric, 2–24 cm, axis glandular-pubescent, verticillasters 2–7, cymes 3–11-flowered, 2 per node;

proximal bracts triangular to lanceolate, 6–50 × 1–22 mm, margins entire or ± serrate;

peduncles and pedicels ascending to erect, glandular-pubescent.

Flowers

calyx lobes ovate, 1.6–6 × 1.4–3 mm, margins entire or erose, glabrous;

corolla red to crimson, with reddish violet nectar guides, bilaterally symmetric, weakly bilabiate, tubular, 22–30 mm, densely yellow-villous internally abaxially, tube 6–8 mm, throat 6–8 mm diam., constricted at orifice, rounded abaxially, abaxial lobes spreading to reflexed, adaxial lobes projecting to erect;

stamens included, pollen sacs divergent, 1.5–1.9 mm, sides glabrous, sutures denticulate, teeth to 0.1 mm;

staminode 15–17 mm, included or slightly exserted, tip straight to recurved, distal 1–3 mm sparsely lanate, hairs yellow, to 1.5 mm;

style 18–28 mm, usually exserted.

calyx lobes ovate to ovate-oblong or lanceolate, 4–10 × (1–)1.5–2.5 mm, glandular-pubescent;

corolla lavender to purple or violet, with reddish purple nectar guides, ventricose to ventricose-ampliate, 20–30 mm, glandular-pubescent externally, sparsely to densely white-lanate internally, tube 6–7 mm, throat abruptly inflated, 8–10 mm diam., 2-ridged abaxially;

stamens included, pollen sacs opposite, subexplanate, 1.2–1.5 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate;

staminode 21–26 mm, exserted, 0.3–0.4 mm diam., tip straight to recurved, distal 6–10 mm sparsely to moderately lanate, hairs white or yellow, to 3 mm;

style 16–19 mm.

Capsules

10–15 × 5–9 mm.

5–9 × 3.5–5 mm, glabrous.

2n

= 16.

Penstemon cardinalis

Penstemon rattanii

Distribution
from FNA
NM; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Penstemon cardinalis occurs as two weakly differentiated varieties in isolated ranges of the Southern Rocky Mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Penstemon rattanii comprises two morphologically distinct and geographically allopatric varieties confined to the Coast Range of California and Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx lobes 1.6–3.5(–4) × 1.4–2 mm; cauline leaf blades elliptic to spatulate or ovate, not leathery; proximal bracts ovate to lanceolate.
var. cardinalis
1. Calyx lobes 4–6 × 2.3–3 mm; cauline leaf blades ovate to subcordate, sometimes elliptic, leathery; proximal bracts suborbiculate to ovate.
var. regalis
1. Calyx lobes usually longer than capsules, lanceolate, sometimes ovate, (5.5–)6.5–10 mm, apices acute to acuminate, sometimes obtuse; nw California, sw Oregon.
var. rattanii
1. Calyx lobes usually shorter than capsules, ovate-oblong, 4–7 mm, apices obtuse to acute; wc California.
var. kleei
Source FNA vol. 17, p. 147. FNA vol. 17, p. 219.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Elmigera Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. cardinalis var. cardinalis, P. cardinalis var. regalis
P. rattanii var. kleei, P. rattanii var. rattanii
Name authority Wooton & Standley: Contr. U.S. Natl. Herb. 16: 171. (1913) — (as Pentstemon) A. Gray: Proc. Amer. Acad. Arts 15: 50. (1879) — (as Pentstemon rattani)
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