Appalachian beardtongue, eastern gray beardtongue
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larch-leaf beardtongue
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Herbs. |
Plants cespitose. |
ascending to erect, (20–)35–80 cm, retrorsely hairy and glandular-pubescent, sometimes only glandular-pubescent distally, not glaucous. |
erect, 10–30 cm, glabrous or puberulent. |
basal and cauline, not leathery, sparsely to densely pubescent, rarely glabrate; basal and proximal cauline 28–170 × 11–42 mm, blade orbiculate to obovate or elliptic, base tapered, margins irregularly crenate to irregularly serrate, apex rounded to obtuse; cauline 3–7 pairs, sessile or proximals short-petiolate, 28–170 × 9–40 mm, blade oblanceolate to lanceolate, proximals usually ± lyrate, base clasping or tapered, margins crenate to serrate, apex obtuse to acute. |
basal and cauline, not leathery, glabrous, rarely puberulent proximally, especially along margins; basal and proximal cauline sessile, 15–20(–40) × 0.5–1.5 mm, blade linear, base tapered to ± clasping, margins entire, apex acute; cauline 3–7 pairs, sessile, 11–35 × 0.3–1 mm, blade linear, base tapered to ± clasping, margins entire, apex acute. |
interrupted, conic, (5–)8–23 cm, axis ± pubescent and glandular-pubescent, verticillasters 3–6, cymes 5–13-flowered, 2 per node; proximal bracts lanceolate, 8–60 × 2–20 mm, margins entire or ± crenate to serrate; peduncles and pedicels spreading to ascending, ± pubescent and glandular-pubescent. |
continuous or interrupted, cylindric, 3–15 cm, axis glabrous or retrorsely hairy, verticillasters 3–7, cymes 1- or 2(or 3)-flowered, 1 or 2 per node; proximal bracts linear, 10–30 × 0.3–1 mm; peduncles and pedicels glabrous, rarely puberulent. |
calyx lobes lanceolate, 3.2–6(–7) × 1.4–1.8(–2) mm, sparsely to moderately glandular-pubescent; corolla lavender to purple, with lavender or purple nectar guides, ventricose, 20–30 mm, glandular-pubescent externally, ± white-pubescent internally abaxially, tube (4–)5–8(–10) mm, throat abruptly inflated, 5–8 mm diam., slightly 2-ridged abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, navicular, 1–1.5 mm, dehiscing completely, connective splitting, sides glabrous, rarely sparsely pubescent, hairs white, to 0.2 mm, sutures papillate; staminode (11–)14–22 mm, exserted, 0.5–0.7 mm diam., tip straight to slightly recurved, distal (7–)10–12 mm ± pubescent, hairs yellowish, to 2 mm; style 13–15 mm. |
calyx lobes ovate, 3.5–5.8 × 1.3–2.2 mm, glabrous; corolla white or pink to violet or purple, without nectar guides, tubular-funnelform, 9–18 mm, glabrous externally, moderately yellow-villous internally abaxially, tube 4–5 mm, throat gradually inflated, not constricted, 5–6 mm diam., rounded abaxially; stamens included, pollen sacs opposite, navicular, 1–1.5 mm, dehiscing completely, sutures papillate; staminode 7–11 mm, included or reaching orifice, 0.5–0.7 mm diam., tip straight to recurved, distal 3–5 mm pilose, hairs yellow or yellow-orange, to 1 mm; style 9–12 mm. |
5–7 × 4–5 mm, glabrous. |
4–5 × 2.5–4 mm. |
= 16. |
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Flowering (Apr–)May–Jul(–Sep). |
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Woodlands, thickets, cliffs, barrens. |
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200–1600 m. (700–5200 ft.) |
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AL; GA; IL; IN; KY; MD; NC; OH; PA; SC; TN; VA; WV
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CO; MT; WY
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F. W. Pennell (1935) acknowledged that Penstemon canescens varies greatly in leaf size, flower size, and vestiture. He distinguished subsp. brittoniorum by its glabrate leaves with sharply serrate margins and dark purple corollas. Plants with those features are encountered mostly in the southern part of the range of the species, although they are essentially sympatric with subsp. canescens and sometimes in mixed populations with plants with the features of subsp. canescens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Molecular data (A. D. Wolfe et al. 2006; C. A. Wessinger et al. 2016) place Penstemon laricifolius with or near sect. Cristati. Two varieties, differentiated by corolla color, are difficult to distinguish in the herbarium, where corollas are usually brown. Populations with a mix of white, pink, and purple corollas have been documented in Wyoming (B. L. Heidel and J. Handley 2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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FNA vol. 17, p. 197. |
FNA vol. 17, p. 138. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. laevigatus var. canescens, P. brittoniorum, P. canescens subsp. brittoniorum |
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(Britton) Britton: Mem. Torrey Bot. Club 5: 291. (1894) — (as Pentstemon) |
Hooker & Arnott: Bot. Beechey Voy., 376. (1839) — (as Pentstemon) |
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