Appalachian beardtongue, eastern gray beardtongue
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limestone beardtongue
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Herbs. |
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ascending to erect, (20–)35–80 cm, retrorsely hairy and glandular-pubescent, sometimes only glandular-pubescent distally, not glaucous. |
ascending to erect, 5–35 cm, puberulent. |
basal and cauline, not leathery, sparsely to densely pubescent, rarely glabrate; basal and proximal cauline 28–170 × 11–42 mm, blade orbiculate to obovate or elliptic, base tapered, margins irregularly crenate to irregularly serrate, apex rounded to obtuse; cauline 3–7 pairs, sessile or proximals short-petiolate, 28–170 × 9–40 mm, blade oblanceolate to lanceolate, proximals usually ± lyrate, base clasping or tapered, margins crenate to serrate, apex obtuse to acute. |
basal and cauline, ± leathery, ± retrorsely hairy, sometimes glabrate abaxially; basal and proximal cauline petiolate, 15–55 × 6–25 mm, blade elliptic to broadly ovate, base tapered, rarely truncate, margins entire or remotely and obscurely serrate, apex obtuse to acute; cauline 2 or 3 pairs, sessile, 25–75 × 10–21 mm, blade lanceolate to oblanceolate, rarely ovate, base truncate to subcordate-clasping, margins entire or obscurely dentate, apex obtuse to acute. |
interrupted, conic, (5–)8–23 cm, axis ± pubescent and glandular-pubescent, verticillasters 3–6, cymes 5–13-flowered, 2 per node; proximal bracts lanceolate, 8–60 × 2–20 mm, margins entire or ± crenate to serrate; peduncles and pedicels spreading to ascending, ± pubescent and glandular-pubescent. |
continuous or interrupted, cylindric, 6–18 cm, axis glandular-pubescent, verticillasters 2–8, cymes (1–)3–7-flowered, 2 per node; proximal bracts lanceolate to ovate, 13–38 × 9–20 mm; peduncles and pedicels glandular-pubescent. |
calyx lobes lanceolate, 3.2–6(–7) × 1.4–1.8(–2) mm, sparsely to moderately glandular-pubescent; corolla lavender to purple, with lavender or purple nectar guides, ventricose, 20–30 mm, glandular-pubescent externally, ± white-pubescent internally abaxially, tube (4–)5–8(–10) mm, throat abruptly inflated, 5–8 mm diam., slightly 2-ridged abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, navicular, 1–1.5 mm, dehiscing completely, connective splitting, sides glabrous, rarely sparsely pubescent, hairs white, to 0.2 mm, sutures papillate; staminode (11–)14–22 mm, exserted, 0.5–0.7 mm diam., tip straight to slightly recurved, distal (7–)10–12 mm ± pubescent, hairs yellowish, to 2 mm; style 13–15 mm. |
calyx lobes lanceolate, 5–7.5 × 0.9–1.8 mm, glandular-pubescent; corolla pink to rose red or rose purple, with reddish nectar guides, tubular-funnelform, 12–17 mm, sparsely white-lanate internally abaxially, tube 4–5 mm, throat gradually inflated, not constricted at orifice, 2.5–4 mm diam., slightly 2-ridged abaxially; stamens included, pollen sacs opposite, subexplanate to explanate, 0.6–0.7 mm, dehiscing completely, sutures smooth; staminode 7–9 mm, included, 0.2–0.3 mm diam., tip straight to recurved, distal 5–6 mm densely pilose, hairs golden yellow, to 0.8 mm; style 4–6 mm. |
5–7 × 4–5 mm, glabrous. |
6–9 × 4–5.5 mm. |
= 16. |
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Flowering (Apr–)May–Jul(–Sep). |
Flowering Apr–May. |
Woodlands, thickets, cliffs, barrens. |
Limestone crevices, rocky slopes, pine-juniper woodlands. |
200–1600 m. (700–5200 ft.) |
1200–2000 m. (3900–6600 ft.) |
AL; GA; IL; IN; KY; MD; NC; OH; PA; SC; TN; VA; WV
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CA; NV |
F. W. Pennell (1935) acknowledged that Penstemon canescens varies greatly in leaf size, flower size, and vestiture. He distinguished subsp. brittoniorum by its glabrate leaves with sharply serrate margins and dark purple corollas. Plants with those features are encountered mostly in the southern part of the range of the species, although they are essentially sympatric with subsp. canescens and sometimes in mixed populations with plants with the features of subsp. canescens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon calcareus is known from Inyo and San Bernardino counties, California, and Esmeralda County, Nevada. The later homonym, P. calcareus M. E. Jones, is referable to P. petiolatus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 197. |
FNA vol. 17, p. 131. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. laevigatus var. canescens, P. brittoniorum, P. canescens subsp. brittoniorum |
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(Britton) Britton: Mem. Torrey Bot. Club 5: 291. (1894) — (as Pentstemon) |
Brandegee: Zoë 5: 152. (1903) — (as Pentstemon) |
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