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mat beardtongue, mat penstemon

Palmer's beardtongue, Palmer's penstemon, scented beardtongue, scented penstemon

Habit Herbs.
Stems

prostrate, only tip ascending, 2–8(–10) cm, not cinereous, retrorsely hairy, hairs pointed.

ascending to erect, 43–140 cm, glaucous.

Leaves

not leathery, not cinereous, retrorsely hairy, hairs pointed;

cauline 3–20 pairs, petiolate, 2.5–14(–21) × 1–4 mm, blade obovate to spatulate, oblanceolate, or linear, base tapered, apex rounded to acute or mucronate.

basal and proximal cauline (28–)52–120(–145) × (5–)25–50(–60) mm, blade ovate, base tapered, margins coarsely dentate, apex obtuse to acute;

cauline 5–8 pairs, sessile or connate-perfoliate, 25–130 × 7–45 mm, blade ovate to triangular, base of distals auriculate-clasping or connate-perfoliate, margins coarsely dentate, distals sometimes entire, apex acute.

Thyrses

continuous, cylindric to secund, 0.5–3 cm, axis retrorsely hairy to retrorsely cinereous, hairs pointed, sometimes also glandular-pubescent, verticillasters 1–5, cymes 1–3-flowered, 1 per node;

proximal bracts linear, 2.8–13(–17) × 0.7–3 mm;

peduncles and pedicels ascending to erect, retrorsely hairy to retrorsely cinereous, hairs pointed, and glandular-pubescent.

interrupted, rarely continuous, secund, 7–63 cm, axis glabrous or glandular-pubescent, verticillasters (4–)7–17, cymes (1 or)2–5-flowered;

proximal bracts ovate to lanceolate, 8–40 × 4–38 mm;

peduncles and pedicels ascending to erect, glabrous or glandular-pubescent.

Flowers

calyx lobes lanceolate to linear, 4–6.5(–8.5) × 1–1.5(–1.9) mm, herbaceous- or narrowly scarious-margined, retrorsely hairy, hairs pointed, and sparsely glandular-pubescent;

corolla blue to purplish lavender or lavender, lined internally abaxially with reddish violet nectar guides, funnelform, 10–17(–21) mm, yellow-villous internally abaxially, tube 5–6 mm, throat gradually inflated, 4.5–6 mm diam., 2-ridged abaxially;

stamens: longer pair reaching orifice or exserted, filaments glabrous, pollen sacs opposite, navicular to explanate, 0.6–1.2 mm, sutures papillate;

staminode 9–12 mm, reaching orifice, flattened distally, 0.3–0.5 mm diam., tip straight to recurved, distal 7–9 mm densely pilose, hairs golden yellow, to 1 mm;

style 13–15 mm.

calyx lobes ovate, (3.4–)4.2–6(–7.5) × 1.9–3.6 mm, glabrous or glandular-pubescent;

corolla pale pink to white or pinkish lavender, with reddish purple nectar guides, strongly bilabiate, ventricose-ampliate, 25–38 mm, glandular-pubescent externally, glandular-pubescent internally, sometimes also sparsely white-lanate abaxially, tube 4–8 mm, length 1–1.4 times calyx lobes, throat abruptly inflated, constricted at orifice, 12–21 mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen sacs navicular, 1.6–2.4 mm, sutures smooth;

staminode 25–30 mm, exserted, 0.9–1.1 mm diam., tip strongly recurved to coiled, distal 8–10 mm, and especially tip, moderately to densely lanate, hairs yellow, to 3.5 mm, proximal 6–8 mm glandular-puberulent;

style 19–23 mm, glabrous or sparsely glandular-pubescent proximally.

Capsules

3.5–5 × 3–4 mm.

11–16 × 6–8 mm, sparsely glandular-pubescent distally.

Penstemon caespitosus

Penstemon palmeri

Distribution
from FNA
AZ; CO; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; NM; NV; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

The varieties of Penstemon caespitosus are weakly differentiated and largely allopatric.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Penstemon ×bryantiae D. D. Keck, a hybrid between P. palmeri and P. spectabilis, was described based on plants that volunteered in the Rancho Santa Ana Botanic Garden in 1935 (D. D. Keck 1937b). Penstemon ×mirus A. Nelson, a putative hybrid between P. eatonii and P. palmeri, has been reported from Arizona (A. Nelson 1938).

A poultice prepared from Penstemon palmeri is applied to snakebites by the Kayenta Navajo of northeastern Arizona (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pollen sacs 0.9–1.2 mm; leaf blades oblanceolate; Arizona, Utah.
var. desertipicti
1. Pollen sacs 0.6–0.9 mm; leaf blades obovate, spatulate, oblanceolate, or linear; Colorado, Utah, Wyoming.
→ 2
2. Leaf blades oblanceolate to linear; pollen sacs explanate, 0.6–0.8 mm; Colorado, Utah, Wyoming.
var. caespitosus
2. Leaf blades obovate or spatulate; pollen sacs navicular to subexplanate, 0.8–0.9 mm; Colorado, Utah.
var. perbrevis
1. Corolla tubes 7–8 mm; distal cauline leaves sessile, sometimes connate-perfoliate.
var. macranthus
1. Corolla tubes 4–6 mm; distal cauline leaves connate-perfoliate.
→ 2
2. Peduncles, pedicels, and calyx lobes glandular-pubescent; pollen sacs 1.8–2.4 mm.
var. palmeri
2. Peduncles, pedicels, and calyx lobes glabrous; pollen sacs 1.6–2(–2.2) mm.
var. eglandulosus
Source FNA vol. 17, p. 100. FNA vol. 17, p. 252.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Caespitosi Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. caespitosus var. caespitosus, P. caespitosus var. desertipicti, P. caespitosus var. perbrevis
P. palmeri var. eglandulosus, P. palmeri var. macranthus, P. palmeri var. palmeri
Name authority Nuttall ex A. Gray: Proc. Amer. Acad. Arts 6: 66. (1862) — (as Pentstemon) A. Gray: Proc. Amer. Acad. Arts 7: 379. (1868) — (as Pentstemon)
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