azure beardtongue, azure penstemon, blue penstemon
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Penland's beardtongue
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Subshrubs. |
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ascending to erect, 13–50(–70) cm, glabrous, glaucous. |
ascending to erect, 7–25 cm, retrorsely hairy or puberulent, not glaucous. |
basal and cauline, basal sometimes few, opposite, glabrous, glaucous; basal and proximal cauline 15–60 × 2–10(–20) mm, blade obovate to oblanceolate or linear, base tapered, margins entire, apex obtuse to acute; cauline 3–20 pairs, sessile or proximals short-petiolate, 10–90 × 2–20 mm, blade elliptic to ovate, lanceolate, or linear, base tapered to clasping or cordate-clasping, margins entire, apex obtuse to acute. |
basal and cauline, not leathery, puberulent or retrorsely hairy, not glaucous; basal and proximal cauline 15–80 × 0.8–2 mm, blade linear, base tapered, margins entire, apex obtuse to acute; cauline 1–3 pairs, sessile, 10–75 × 0.5–1.5 mm, blade linear, base tapered, apex acute. |
interrupted or continuous, secund, (3–)6–31 cm, axis glabrous, verticillasters 3–10, cymes 1–3(or 4)-flowered, 2 per node; proximal bracts ovate to lanceolate, 4–50 × 1–32 mm; peduncles and pedicels ascending to erect, glabrous. |
continuous, secund, 3–10 cm, axis glandular-pubescent, verticillasters 2–6, cymes 1- or 2(–4)-flowered, 2 per node; proximal bracts linear, 10–56 × 0.5–1.5 mm; peduncles and pedicels glandular-pubescent. |
calyx lobes ovate to oblong or obovate, 2.5–6.5 × 1.4–3.2 mm, glabrous; corolla violet to blue or purple, with or without purple nectar guides, funnelform to ventricose, 18–35 mm, glabrous externally, glabrous internally, tube 7–9 mm, throat gradually inflated, 8–10 mm diam., slightly 2-ridged abaxially; stamens: longer pair reaching orifice or exserted, filaments glabrous, pollen sacs parallel, 1.8–3.5 mm, distal 1/3–1/2 indehiscent, sides hispidulous to pilose, hairs white, to 0.8 mm near filament attachment, sutures denticulate, teeth to 0.3 mm; staminode 12–19 mm, included, 0.4–0.6 mm diam., glabrous; style 16–22 mm. |
calyx lobes lanceolate, 4.5–5.5 × 1.8–2.5 mm, glabrous to sparsely glandular; corolla blue to bluish violet, with reddish purple nectar guides, funnelform, 13–16 mm, glabrous externally, glabrous internally, tube 5–6 mm, throat gradually inflated, not constricted at orifice, 4.5–5.5 mm diam., rounded abaxially; stamens included, pollen sacs divergent to opposite, navicular, 1.1–1.5 mm, dehiscing incompletely, connective not splitting, sides moderately hirsute, hairs white, to 0.5 mm, sutures denticulate, teeth to 0.1 mm; staminode 8–9 mm, included, 0.9–1.1 mm diam., tip straight, distal 6–7 mm densely pubescent, hairs yellowish orange, to 0.9 mm; style 8–10 mm. |
7–11 × 14–6 mm. |
6–9(–14) × 4–6 mm. |
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Flowering Jun–Jul. |
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Clayey soils, sagebrush shrublands. |
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2300–2400 m. (7500–7900 ft.) |
CA; OR
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CO |
Varieties 2 (2 in the flora). Penstemon azureus occurs from southwestern Oregon to central California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon penlandii is known from Middle Park in Grand County. Plants grow on strongly seleniferous shales of the Troublesome Formation (S. L. O’Kane 1988). The species is listed as endangered by the U.S. Department of the Interior. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Calyx lobes obovate to oblong, 2.5–4.8 × 2–3.2 mm, apices cuspidate to mucronate; cauline leaves 10–65 × 4–20 mm, blades elliptic to ovate or lanceolate. | var. azureus |
1. Calyx lobes ovate, 5–6.5 × 1.4–2.3 mm, apices acuminate to caudate; cauline leaves 40–90 × 2–6 mm, blades lanceolate to linear. | var. angustissimus |
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FNA vol. 17, p. 231. |
FNA vol. 17, p. 176. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Bentham: Pl. Hartw., 327. (1849) — (as Pentstemon) |
W. A. Weber: Phytologia 60: 459. (1986) |
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