azure beardtongue, azure penstemon, blue penstemon
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blowout beardtongue
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Subshrubs. |
Plants: caudex rhizomelike. |
ascending to erect, 13–50(–70) cm, glabrous, glaucous. |
decumbent to ascending, (15–)20–48 cm, glabrous. |
basal and cauline, basal sometimes few, opposite, glabrous, glaucous; basal and proximal cauline 15–60 × 2–10(–20) mm, blade obovate to oblanceolate or linear, base tapered, margins entire, apex obtuse to acute; cauline 3–20 pairs, sessile or proximals short-petiolate, 10–90 × 2–20 mm, blade elliptic to ovate, lanceolate, or linear, base tapered to clasping or cordate-clasping, margins entire, apex obtuse to acute. |
essentially cauline, basal absent or reduced, glabrous; basal and proximal cauline (25–)55–130(–175) × 3–25 mm, blade linear to lanceolate, base tapered, apex acute to acuminate; cauline 2–10 pairs, sessile, 60–110(–120) × 7–30 mm, blade lanceolate to linear, base clasping, apex acuminate to long-acuminate. |
interrupted or continuous, secund, (3–)6–31 cm, axis glabrous, verticillasters 3–10, cymes 1–3(or 4)-flowered, 2 per node; proximal bracts ovate to lanceolate, 4–50 × 1–32 mm; peduncles and pedicels ascending to erect, glabrous. |
continuous, cylindric, (2–)6–21(–34) cm, axis glabrous, verticillasters (2–)6–10(–17), cymes 1–8-flowered; proximal bracts ovate, 58–120 × 20–45 mm; peduncles and pedicels glabrous. |
calyx lobes ovate to oblong or obovate, 2.5–6.5 × 1.4–3.2 mm, glabrous; corolla violet to blue or purple, with or without purple nectar guides, funnelform to ventricose, 18–35 mm, glabrous externally, glabrous internally, tube 7–9 mm, throat gradually inflated, 8–10 mm diam., slightly 2-ridged abaxially; stamens: longer pair reaching orifice or exserted, filaments glabrous, pollen sacs parallel, 1.8–3.5 mm, distal 1/3–1/2 indehiscent, sides hispidulous to pilose, hairs white, to 0.8 mm near filament attachment, sutures denticulate, teeth to 0.3 mm; staminode 12–19 mm, included, 0.4–0.6 mm diam., glabrous; style 16–22 mm. |
calyx lobes lanceolate to linear, 8–13 × 1–3 mm, margins entire or erose, herbaceous or scarious, glabrous; corolla lavender to bluish, usually with magenta nectar guides, ampliate, 21–28 mm, glabrous externally, glabrous internally, tube 7–9 mm, throat abruptly inflated, 9–11 mm diam., rounded abaxially; stamens included, pollen sacs opposite, 1.1–2 mm, sutures papillate; staminode 13–16 mm, included, 1.2–3 mm diam., tip recurved, distal 0.1–5 mm sparsely to densely villous, hairs golden yellow, to 1 mm, rarely glabrous; style 13–23 mm. |
7–11 × 14–6 mm. |
8–12 × 5–9 mm. |
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= 16. |
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Flowering May–Jul(–Sep). |
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Blowouts in sand dunes. |
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900–2300 m. (3000–7500 ft.) |
CA; OR
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NE; WY |
Varieties 2 (2 in the flora). Penstemon azureus occurs from southwestern Oregon to central California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon haydenii is known from the Nebraska Sandhills, where extant populations occur in Box Butte, Cherry, Garden, Hooker, Morrill, and Sheridan counties. Historic populations occurred in Thomas County (D. M. Sutherland 1988). Penstemon haydenii was discovered in northern Carbon County, Wyoming, in the late 1990s some 300 km west of the nearest Nebraska populations; it might have been collected in Wyoming as early as 1877 (W. Fertig 2001). A morphometric analysis of Nebraska and Wyoming plants revealed differences that could justify recognition of Wyoming populations as a distinct variety (C. C. Freeman 2015). It is listed as endangered by the U.S. Department of the Interior. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Calyx lobes obovate to oblong, 2.5–4.8 × 2–3.2 mm, apices cuspidate to mucronate; cauline leaves 10–65 × 4–20 mm, blades elliptic to ovate or lanceolate. | var. azureus |
1. Calyx lobes ovate, 5–6.5 × 1.4–2.3 mm, apices acuminate to caudate; cauline leaves 40–90 × 2–6 mm, blades lanceolate to linear. | var. angustissimus |
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FNA vol. 17, p. 231. |
FNA vol. 17, p. 119. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Bentham: Pl. Hartw., 327. (1849) — (as Pentstemon) |
S. Watson: Bot. Gaz. 16: 311. (1891) — (as Pentstemon haydeni) |
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