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azure beardtongue, azure penstemon, blue penstemon

white beardtongue, white penstemon

Habit Subshrubs.
Stems

ascending to erect, 13–50(–70) cm, glabrous, glaucous.

ascending to erect, (10–)15–50(–55) cm, retrorsely hairy proximally, glandular-pubescent distally.

Leaves

basal and cauline, basal sometimes few, opposite, glabrous, glaucous;

basal and proximal cauline 15–60 × 2–10(–20) mm, blade obovate to oblanceolate or linear, base tapered, margins entire, apex obtuse to acute;

cauline 3–20 pairs, sessile or proximals short-petiolate, 10–90 × 2–20 mm, blade elliptic to ovate, lanceolate, or linear, base tapered to clasping or cordate-clasping, margins entire, apex obtuse to acute.

basal and cauline, not leathery, glabrate or puberulent to scabrous;

basal and proximal cauline petiolate, 20–85(–110) × (4–)7–18(–20) mm, blade oblanceolate or obovate to lanceolate, base tapered, margins entire to obscurely or distinctly serrate, apex obtuse to acute;

cauline 2–5(or 6) pairs, sessile or proximals short-petiolate, 25–65 × (3–)7–19(–21) mm, blade ovate to lanceolate, base tapered to clasping, margins entire or serrate to dentate, apex acute.

Thyrses

interrupted or continuous, secund, (3–)6–31 cm, axis glabrous, verticillasters 3–10, cymes 1–3(or 4)-flowered, 2 per node;

proximal bracts ovate to lanceolate, 4–50 × 1–32 mm;

peduncles and pedicels ascending to erect, glabrous.

continuous or interrupted, cylindric, 4–24(–30) cm, axis densely glandular-pubescent, verticillasters (2 or)3–10, cymes 2–7-flowered, 2 per node;

proximal bracts lanceolate, 17–65 × 3–17 mm;

peduncles and pedicels densely glandular-pubescent.

Flowers

calyx lobes ovate to oblong or obovate, 2.5–6.5 × 1.4–3.2 mm, glabrous;

corolla violet to blue or purple, with or without purple nectar guides, funnelform to ventricose, 18–35 mm, glabrous externally, glabrous internally, tube 7–9 mm, throat gradually inflated, 8–10 mm diam., slightly 2-ridged abaxially;

stamens: longer pair reaching orifice or exserted, filaments glabrous, pollen sacs parallel, 1.8–3.5 mm, distal 1/3–1/2 indehiscent, sides hispidulous to pilose, hairs white, to 0.8 mm near filament attachment, sutures denticulate, teeth to 0.3 mm;

staminode 12–19 mm, included, 0.4–0.6 mm diam., glabrous;

style 16–22 mm.

calyx lobes ovate to lanceolate, 4–7 × 1.5–3 mm, glandular-pubescent;

corolla white, rarely tinged pink or lavender, with red or reddish purple nectar guides, funnelform, (12–)16–20 mm, glandular-pubescent internally, tube 4–6 mm, throat gradually inflated, not constricted at orifice, (4–)6–8 mm diam., rounded abaxially;

stamens included, pollen sacs opposite, explanate, 0.7–1.1 mm, dehiscing completely, sutures smooth;

staminode 8–9 mm, included, 0.3–0.4 mm diam., tip straight to recurved, distal 3–6 mm sparsely to moderately villous, hairs yellowish, to 1 mm;

style 9–11(–13) mm.

Capsules

7–11 × 14–6 mm.

8–12 × 4–7 mm.

2n

= 16.

Penstemon azureus

Penstemon albidus

Phenology Flowering Apr–Jul(–Sep).
Habitat Silty or gravelly soils, mixed-grass and shortgrass prairies.
Elevation 300–1800 m. (1000–5900 ft.)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; IA; KS; MN; MT; ND; NE; NM; OK; SD; TX; WY; AB; MB; ON; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Penstemon azureus occurs from southwestern Oregon to central California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon albidus is remarkably uniform throughout its range. It probably is introduced in Ontario.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx lobes obovate to oblong, 2.5–4.8 × 2–3.2 mm, apices cuspidate to mucronate; cauline leaves 10–65 × 4–20 mm, blades elliptic to ovate or lanceolate.
var. azureus
1. Calyx lobes ovate, 5–6.5 × 1.4–2.3 mm, apices acuminate to caudate; cauline leaves 40–90 × 2–6 mm, blades lanceolate to linear.
var. angustissimus
Source FNA vol. 17, p. 231. FNA vol. 17, p. 128.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. azureus var. angustissimus, P. azureus var. azureus
Name authority Bentham: Pl. Hartw., 327. (1849) — (as Pentstemon) Nuttall: Gen. N. Amer. Pl. 2: 53. (1818) — (as Pentstemon albidum)
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