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Eustis Lake beardtongue, southern beardtongue

Ward's beardtongue

Habit Herbs.
Stems

erect, 30–86 cm, retrorsely hairy and glandular-lanate, not glaucous.

ascending to erect, (6–)15–42 cm, ± retrorsely hairy or pubescent, not glaucous.

Leaves

basal and cauline, not leathery, retrorsely hairy, sometimes also glandular-pubescent, rarely glabrate;

basal and proximal cauline 32–130 × 7–40 mm, blade spatulate to oblanceolate, base tapered, margins entire or ± serrate, apex rounded to obtuse;

cauline 5–8 pairs, sessile or proximals short-petiolate, 16–122 × 3–23 mm, blade lanceolate, base clasping or tapered, margins ± serrate, apex acute to acuminate.

basal and cauline, ± leathery or not, densely retrorsely hairy, not glaucous;

basal and proximal cauline 20–70 × 3–16(–22) mm, blade oblanceolate, base tapered, margins entire, apex obtuse to acute;

cauline 2–5 pairs, short-petiolate or sessile, 19–72 × 3–9 mm, blade oblanceolate to lanceolate, base tapered to truncate, apex obtuse to acute.

Thyrses

interrupted, narrowly conic, rarely conic, 7–26 cm, axis ± glandular-pubescent, verticillasters 3–6, cymes 2–6-flowered, usually only 1 branch of each cyme elongating, others nearly sessile or distinctly shorter, 2 per node;

proximal bracts lanceolate, 3–20 × 1–4 mm, margins entire;

peduncles and pedicels erect, sometimes ascending, ± glandular-pubescent.

continuous, secund, (4–)10–22 cm, axis retrorsely hairy to retrorsely pubescent or pubescent, verticillasters (3–)5–8, cymes 1–5-flowered, 2 per node;

proximal bracts oblanceolate to lanceolate, 25–70 × 3–9 mm;

peduncles and pedicels retrorsely hairy and glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, 4–5.2 × 1.6–2.3 mm, sparsely glandular-pubescent;

corolla white to pinkish or light lavender, with dark purple nectar guides, tubular, 20–25 mm, glandular-pubescent externally, moderately white-lanate internally abaxially, tube 4–5 mm, throat abruptly inflated, 4.5–6 mm diam., 2-ridged abaxially;

stamens included, pollen sacs opposite, navicular, 1.2–1.5 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate;

staminode 11–14 mm, exserted, 0.4–0.7 mm diam., tip straight to slightly recurved, distal 8–10 mm densely pilose, hairs yellow, to 2 mm;

style 10–12 mm.

calyx lobes ovate, 6–8.5 × 2.8–3.8 mm, glandular-pubescent;

corolla blue to lavender, with or without faint lavender nectar guides, ventricose, 24–30 mm, glandular-pubescent externally, glabrous internally, tube 8–12 mm, throat gradually inflated, slightly constricted at orifice, 6–9(–12) mm diam., slightly 2-ridged abaxially;

stamens: longer pair reaching orifice or slightly exserted, pollen sacs divergent, sigmoid, 2–2.5 mm, dehiscing incompletely, proximal 1/5–1/4 indehiscent, connective not splitting, sides glabrous, sutures denticulate, teeth to 0.1 mm;

staminode 16–18 mm, included, 0.5–0.8 mm diam., tip straight, glabrous;

style 20–24 mm.

Capsules

6–9 × 4.5–6 mm, glabrous.

12–15 × 8–10 mm.

Penstemon australis

Penstemon wardii

Phenology Flowering Mar–Jun. Flowering May–Jul.
Habitat Sandy pine and oak woodlands, pine savannas, hammocks, granite hills, sandy open areas. Clayey or loamy soils, foothills, desert shrub communities, pinyon-juniper woodlands.
Elevation 10–200 m. (0–700 ft.) 1500–2000 m. (4900–6600 ft.)
Distribution
from FNA
AL; FL; GA; MS; NC; SC; TN; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Penstemon australis is known from the southern Atlantic Coastal Plain and eastern Gulf Coastal Plain. The species is confused most often with P. laxiflorus, which occurs farther west. Penstemon australis is distinguished from P. laxiflorus most readily by the stem vestiture; P. australis usually has stems with a mix of short, eglandular hairs and much longer, glandular hairs, while P. laxiflorus has stems with only short, retrorse hairs or, if glandular hairs also are present, they are sparse and occur just below the inflorescences. Penstemon australis also tends to have narrower inflorescences due to the unequal elongation of cyme branches. Ranges of the two species overlap in southwestern Alabama, and many specimens from Baldwin, Escambia, and Mobile counties are morphologically intermediate.

Many specimens of Penstemon from the Appalachian Piedmont, especially in northeastern Alabama, northern Georgia, central North Carolina, and western South Carolina, combine morphological features of P. australis and P. canescens. These plants usually have equally developed cyme branches (as in P. canescens) but ascending or erect peduncles and pedicels (as in P. australis). Leaf shape generally is intermediate between P. australis and P. canescens. F. W. Pennell in the 1930s annotated most such specimens as P. australis. F. S. Crosswhite (1965e) also included these specimens in his concept of P. australis, assigning them subspecies status, though he never published the subspecies names that he used in his thesis and on herbarium sheets. The relationship of these plants remains unclear; they will key to P. canescens or P. laxiflorus.

A. E. Radford et al. (1968) gave a chromosome number of n = 8 for Penstemon australis; this does not appear to be supported by any other published chromosome count.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon wardii is known from the foothills surrounding Sevier Valley in Millard, Piute, Sanpete, and Sevier counties (E. C. Neese and N. D. Atwood 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 195. FNA vol. 17, p. 183.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority Small: Fl. S.E. U.S., 1060, 1337. (1903) — (as Pentstemon) A. Gray: Proc. Amer. Acad. Arts 12: 82. (1876) — (as Pentstemon wardi)
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