Colorado beardtongue
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Alberta beardtongue, Alberta penstemon
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Herbs. |
ascending to erect, 10–30(–35) cm, retrorsely hairy proximally, glandular-pubescent distally. |
ascending to erect, (10–)15–38 cm, glabrous or sparsely retrorsely hairy, sometimes also sparsely glandular-pubescent. |
basal and cauline, not leathery, glabrous, puberulent, or retrorsely hairy; basal and proximal cauline petiolate, (15–)30–60(–100) × (1–)2–5(–7) mm, blade lanceolate to linear, base tapered, margins entire, apex obtuse to acute; cauline 2–6 pairs, sessile, (25–)40–80 × 2–7 mm, blade lanceolate to linear, base tapered to slightly clasping, margins entire or obscurely denticulate, apex acute to acuminate. |
basal and cauline, ± leathery or not, glabrous, sometimes sparsely retrorsely hairy along midvein abaxially, not glaucous; basal and proximal cauline (13–)40–90 × (3–)6–18 mm, blade ovate to oblanceolate or lanceolate, base tapered, margins ± serrate or dentate, apex obtuse to acute; cauline 2–6 pairs, short-petiolate or sessile, 15–55 × 3–10 mm, blade lanceolate, proximals sometimes oblanceolate to oblong, base tapered or truncate, margins entire or ± serrate to ± dentate, apex obtuse to acute. |
continuous or interrupted, cylindric or slightly secund, (5–)7–23 cm, axis glandular-pubescent, verticillasters 3–8, cymes 2–4-flowered, 2 per node; proximal bracts lanceolate to linear, 18–100 × 2–12 mm; peduncles and pedicels glandular-pubescent. |
continuous or ± interrupted, cylindric, 4–19 cm, axis glabrous or ± glandular-pubescent, verticillasters 4–8, cymes (1 or)2–6-flowered, 2 per node; proximal bracts lanceolate, 9–40 × 3–15 mm, margins entire or ± serrate to dentate; peduncles and pedicels erect, sometimes ascending, glandular-pubescent. |
calyx lobes ovate to lanceolate, (6–)7–9 × 1–2 mm, densely glandular-pubescent; corolla violet to lilac or pinkish blue, with magenta or violet nectar guides, weakly bilabiate, tubular-funnelform, (16–)18–22(–24) mm, moderately white-pilose internally abaxially, tube 4–6 mm, throat gradually inflated, not constricted at orifice, 7–9 mm diam., rounded abaxially; stamens included, pollen sacs divergent, navicular, 1.2–1.5 mm, dehiscing incompletely, connective not splitting, sutures papillate; staminode 13–16 mm, reaching orifice or exserted, 0.6–0.9 mm diam., tip recurved, distal 10–15 mm moderately to densely villous, hairs orangish, to 2.5 mm; style 11–14 mm. |
calyx lobes ovate, 2.8–4.8 × 0.9–1.8 mm, glandular-pubescent; corolla blue to light blue or violet, with reddish purple nectar guides, funnelform, 13–20 mm, glandular-pubescent externally, sparsely white-villous internally abaxially, tube 4–5 mm, throat gradually inflated, 4–5 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, subexplanate to explanate, 0.5–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate; staminode 8–9 mm, included, 0.2–0.3 mm diam., tip straight to recurved, distal 0.5–2 mm ± pilose, hairs golden yellow, to 0.8 mm; style 9–11 mm. |
(6–)8–10 × 3–4 mm. |
4–6 × 3–4 mm, glabrous. |
= 16. |
= 16. |
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Flowering May–Aug. |
Flowering May–Aug. |
Silty or gravelly soils, shortgrass prairies, sagebrush shrublands. |
Gravelly slopes, cliffs, rocky clearings, roadcuts in coniferous forests. |
1200–2500 m. (3900–8200 ft.) |
800–2800 m. (2600–9200 ft.) |
CO; NM
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ID; MT; AB; BC
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Penstemon auriberbis is known from Chaffee, Costilla, Custer, El Paso, Elbert, Fremont, Huerfano, Kiowa, Las Animas, Otero, and Pueblo counties, Colorado, and Colfax and Union counties, New Mexico. The species is unique in sect. Cristati in having incompletely dehiscing anthers. Penstemon parviflorus is known only from the type locality in Montezuma County, Colorado (S. L. O’Kane 1988). A report for McKinley County, New Mexico, has not been confirmed. The type was collected July 1890 by Alice Eastwood, reputedly near Mancos, Colorado. F. W. Pennell (1920b) distinguished P. parviflorus from the more eastern P. auriberbis primarily by the former’s shorter corollas, longer calyx lobes, and more slender stems. D. D. Keck (1938) observed that the type specimen has whorled cauline leaves, which Pennell did not mention in his description, and Keck suggested that P. parviflorus might be a genetically aberrant individual of P. auriberbis that was accidentally introduced into Montezuma County. Whorled cauline leaves are not uncommon in P. auriberbis, and they occur rarely in other species in sect. Cristati. Penstemon parviflorus appears to be P. auriberbis, and the type specimen actually may have come from south-central Colorado; Eastwood collected P. auriberbis in Pueblo County in 1891. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon caelestinus has been treated as a synonym of P. albertinus (D. D. Keck 1945; A. Cronquist 1959). D. V. Clark (1971) grew these and related species in transplant gardens and considered P. caelestinus to be distinct based on flower color, leaf shape and color, phenology, and habitat. A broad zone of hybridization between P. albertinus and P. wilcoxii exists in east-central Idaho and western Montana, especially in habitats disturbed by humans (Clark). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 129. |
FNA vol. 17, p. 190. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. parviflorus |
P. caelestinus |
Pennell: Contr. U.S. Natl. Herb. 20: 339. (1920) |
Greene: Leafl. Bot. Observ. Crit. 1: 167. (1906) — (as Pentstemon) |
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