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stiff-leaf beardtongue, stiffleaf penstemon

Berry's penstemon, mountain pride, Newberry's beardtongue

Habit Herbs. Subshrubs, cespitose.
Stems

ascending to erect, 6–20(–25) cm, retrorsely hairy proximally, glabrous or retrorsely hairy distally, not glaucous.

ascending to erect, (5–)10–32 cm, glabrous or retrorsely hairy, not glaucous.

Leaves

basal and cauline, not leathery, glabrous or ± puberulent;

basal and proximal cauline 15–65 × 1–5(–7) mm, blade narrowly oblanceolate to linear, base tapered, margins entire, apex obtuse to acute;

cauline 2–5 pairs, short-petiolate or sessile, 7–33 × 1–3 mm, blade lanceolate to linear, base sessile to clasping, margins entire, apex obtuse to acute.

persistent, 4–9 pairs, distals distinctly to slightly smaller than proximals, short-petiolate or sessile, (5–)9–42 × 4–14(–18) mm, blade elliptic to obovate, base tapered, cuneate, or cordate-clasping, margins ± serrulate or serrulate-denticulate to serrate, apex rounded to obtuse or acute, glabrous, glaucous.

Thyrses

interrupted, cylindric, 2–11 cm, axis glandular-pubescent, sometimes also retrorsely hairy, verticillasters 2–6, cymes 1–5-flowered, 2 per node;

proximal bracts lanceolate to linear, 9–32 × 1–6 mm, margins entire;

peduncles and pedicels erect, ± glandular-pubescent.

continuous, ± secund, 2–20 cm, axis glandular-pubescent, verticillasters 4–12, cymes 1-flowered;

proximal bracts ovate to lanceolate, 4–11(–20) × 2–8(–15) mm;

peduncles and pedicels ascending, glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, 2.2–4.6 × 1–2.4 mm, glandular-pubescent;

corolla blue to violet, with reddish purple guides, funnelform, 11–17 mm, glandular-pubescent externally, moderately whitish lanate internally abaxially, tube 3–4 mm, throat gradually inflated, 4–6 mm diam., 2-ridged abaxially;

stamens included, pollen sacs divergent to opposite, subexplanate to explanate, 0.6–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate;

staminode 7–8 mm, reaching orifice, 0.2–0.4 mm diam., tip recurved, distal 0.5–1 mm densely pilose, hairs golden yellow, to 0.8 mm;

style 9–10 mm.

calyx lobes lanceolate, 6–10 × 2–2.5 mm, glandular-pubescent;

corolla pink, red, scarlet, lavender, or purple, essentially unlined internally but abaxial ridges sometimes pink to whitish, not personate, funnelform, 22–35 mm, glabrous externally, moderately to densely white-lanate internally abaxially, tube 6–9 mm, throat 5–12 mm diam.;

stamens included or longer pair exserted, pollen sacs 0.8–1.7 mm;

staminode 8–14 mm, slightly flattened distally, 0.1–0.3 mm diam., tip straight, distal 2–8 mm pubescent to lanate, hairs white or yellowish, to 1 mm;

style 20–24 mm.

Capsules

5–7 × 4–5 mm, glabrous.

(8–)10–13 × (4–)5–8 mm.

Penstemon aridus

Penstemon newberryi

Phenology Flowering Jun–Jul.
Habitat Rocky hillsides, sagebrush shrublands, montane grasslands.
Elevation 1500–2900 m. (4900–9500 ft.)
Distribution
from FNA
ID; MT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon aridus is known from the Rocky Mountains in eastern Idaho and southwestern and south-central Montana and the Big Horn and northern Absaroka Mountains of Wyoming. The sometimes grasslike basal leaves distinguish P. aridus from P. humilis and P. virens. At lower elevations or in shaded, forest habitats, plants sometimes have broader basal leaves and tend to be much taller than typical plants; the broadly scarious-margined calyx lobes of P. aridus distinguish it from similar species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Penstemon newberryi is known from the Sierra Nevada and Cascade ranges of California, Nevada, and Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas pink to lavender or purple, 27–35 mm; stamens included, pollen sacs 1.2–1.7 mm.
var. berryi
1. Corollas red, scarlet, or purple, 22–30 mm; stamens: longer pair exserted, pollen sacs 0.8–1.3 mm.
→ 2
2. Corollas red to scarlet; distal leaves distinctly smaller than proximals.
var. newberryi
2. Corollas purple; distal leaves slightly smaller than proximals.
var. sonomensis
Source FNA vol. 17, p. 192. FNA vol. 17, p. 90.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon Plantaginaceae > Penstemon > subg. Dasanthera > sect. Erianthera
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. newberryi var. berryi, P. newberryi var. newberryi, P. newberryi var. sonomensis
Name authority Rydberg: Mem. New York Bot. Gard. 1: 348. (1900) — (as Pentstemon) A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 6(3): 82, plate 14. (1858) — (as Pentstemon)
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