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Penstemon anguineus

Siskiyou beardtongue, tongue leaf penstemon

blue-stem beardtongue, bluestem penstemon

Habit Herbs.
Stems

ascending to erect, (10–)30–80 cm, glabrous, not glaucous.

ascending to erect, (16–)20–60 cm, glabrous or ± puberulent proximally, ± glaucous.

Leaves

basal and cauline, not leathery, glabrous;

basal and proximal cauline 30–150 × 6–40 mm, blade ovate to elliptic or oval, base tapered, margins serrate to dentate, sometimes denticulate or entire, apex obtuse to acute;

cauline 2–4 pairs, sessile or proximals short-petiolate, 11–90 × 5–50 mm, blade oblanceolate to oblong or triangular-ovate, base tapered to cordate-clasping, margins serrate to denticulate, rarely entire, apex obtuse to acute.

basal and cauline, ± leathery or not, glabrous or proximals puberulent proximally or along midveins, ± glaucous;

basal and proximal cauline (30–)50–95 × 10–15(–28) mm, blade spatulate to oblanceolate, base tapered, margins entire, usually undulate, apex rounded to obtuse;

cauline 2–4 pairs, sessile, 20–73 × 4–15 mm, blade lanceolate to elliptic, base truncate to clasping, margins crisped or undulate, apex acute.

Thyrses

interrupted, sometimes continuous, cylindric, (4–)8–40 cm, axis glandular-pubescent, verticillasters 3–10, cymes (2 or)3–15-flowered, 2 per node;

proximal bracts ovate to triangular, 10–85 × 7–40 mm, margins serrate or entire;

peduncles and pedicels ascending, glandular-pubescent.

interrupted, secund to subsecund, (4–)8–31 cm, axis glabrous, verticillasters 5–9(–12), cymes (1 or)2–5-flowered, 2 per node;

proximal bracts ovate to lanceolate, 15–50 × 3–23 mm;

peduncles and pedicels glabrous, peduncles to 18 mm, pedicels 1–7 mm.

Flowers

calyx lobes lanceolate, 4–6.5 × 0.9–2 mm, glandular-pubescent;

corolla light blue to lavender, purple, or violet, with violet nectar guides, ampliate, 13–18 mm, glandular-pubescent externally, glabrous or sparsely white-lanate internally abaxially, tube 5–7 mm, throat gradually inflated, 4–6 mm diam., 2-ridged abaxially;

stamens included, pollen sacs opposite, navicular, 0.8–1.2 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate;

staminode 10–13 mm, exserted, 0.2–0.4 mm diam., tip straight to slightly recurved, distal 2–6 mm glabrous or sparsely pilose, hairs yellow, to 0.6 mm;

style 10–13 mm.

calyx lobes ovate, 3.5–4.5 × 1.4–2 mm, apex acute to acuminate, glabrous;

corolla blue to violet, with violet nectar guides, funnelform to weakly ventricose, 14–18(–20) mm, glabrous or ± glandular externally, glabrous internally, tube 5–8 mm, throat gradually inflated, not constricted at orifice, 5–7 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, pollen sacs divergent, navicular-sigmoid, 1.2–1.7 mm, dehiscing incompletely, proximal 1/4 indehiscent, connective not splitting, sides sparsely to moderately pilose, hairs white, to 0.5(–0.7) mm, sutures denticulate, teeth to 0.1 mm;

staminode: 9–11 mm, reaching orifice, 0.2–0.4 mm diam., tip straight, distal 2–6 mm sparsely pilose, hairs golden yellow, to 0.6 mm;

style 9–12 mm.

Capsules

5–7 × 3–4 mm, glabrous.

6–10 × 4–5 mm.

2n

= 16.

Penstemon anguineus

Penstemon cyanocaulis

Phenology Flowering Jun–Aug. Flowering Apr–Jun.
Habitat Rocky, clayey, or humus soils, openings in coniferous forests. Rocky slopes, pinyon-juniper, oak, and oak-juniper woodlands.
Elevation 600–2000 m. (2000–6600 ft.) 1300–2700 m. (4300–8900 ft.)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon anguineus occurs primarily in the Cascade, Klamath, and North Coast ranges from southwestern Oregon (Curry, Douglas, Jackson, Josephine, and Klamath counties) south into northwestern California (Del Norte, El Dorado, Glenn, Humboldt, Mendocino, Siskiyou, Tehama, and Trinity counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon cyanocaulis is known from Mesa, Montrose, and San Miguel counties, Colorado, and Carbon, Emery, Grand, and San Juan counties, Utah. It has crisped or undulate leaf margins that help distinguish it in the field.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 191. FNA vol. 17, p. 165.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority Eastwood: Bull. Torrey Bot. Club 32: 208. (1905) — (as Pentstemon) Payson: Bot. Gaz. 60: 380. (1915) — (as Pentstemon)
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