Alberta beardtongue, Alberta penstemon
|
Palmer's beardtongue, Palmer's penstemon, scented beardtongue, scented penstemon
|
Herbs. |
Herbs. |
ascending to erect, (10–)15–38 cm, glabrous or sparsely retrorsely hairy, sometimes also sparsely glandular-pubescent. |
ascending to erect, 43–140 cm, glaucous. |
basal and cauline, ± leathery or not, glabrous, sometimes sparsely retrorsely hairy along midvein abaxially, not glaucous; basal and proximal cauline (13–)40–90 × (3–)6–18 mm, blade ovate to oblanceolate or lanceolate, base tapered, margins ± serrate or dentate, apex obtuse to acute; cauline 2–6 pairs, short-petiolate or sessile, 15–55 × 3–10 mm, blade lanceolate, proximals sometimes oblanceolate to oblong, base tapered or truncate, margins entire or ± serrate to ± dentate, apex obtuse to acute. |
basal and proximal cauline (28–)52–120(–145) × (5–)25–50(–60) mm, blade ovate, base tapered, margins coarsely dentate, apex obtuse to acute; cauline 5–8 pairs, sessile or connate-perfoliate, 25–130 × 7–45 mm, blade ovate to triangular, base of distals auriculate-clasping or connate-perfoliate, margins coarsely dentate, distals sometimes entire, apex acute. |
continuous or ± interrupted, cylindric, 4–19 cm, axis glabrous or ± glandular-pubescent, verticillasters 4–8, cymes (1 or)2–6-flowered, 2 per node; proximal bracts lanceolate, 9–40 × 3–15 mm, margins entire or ± serrate to dentate; peduncles and pedicels erect, sometimes ascending, glandular-pubescent. |
interrupted, rarely continuous, secund, 7–63 cm, axis glabrous or glandular-pubescent, verticillasters (4–)7–17, cymes (1 or)2–5-flowered; proximal bracts ovate to lanceolate, 8–40 × 4–38 mm; peduncles and pedicels ascending to erect, glabrous or glandular-pubescent. |
calyx lobes ovate, 2.8–4.8 × 0.9–1.8 mm, glandular-pubescent; corolla blue to light blue or violet, with reddish purple nectar guides, funnelform, 13–20 mm, glandular-pubescent externally, sparsely white-villous internally abaxially, tube 4–5 mm, throat gradually inflated, 4–5 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, subexplanate to explanate, 0.5–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate; staminode 8–9 mm, included, 0.2–0.3 mm diam., tip straight to recurved, distal 0.5–2 mm ± pilose, hairs golden yellow, to 0.8 mm; style 9–11 mm. |
calyx lobes ovate, (3.4–)4.2–6(–7.5) × 1.9–3.6 mm, glabrous or glandular-pubescent; corolla pale pink to white or pinkish lavender, with reddish purple nectar guides, strongly bilabiate, ventricose-ampliate, 25–38 mm, glandular-pubescent externally, glandular-pubescent internally, sometimes also sparsely white-lanate abaxially, tube 4–8 mm, length 1–1.4 times calyx lobes, throat abruptly inflated, constricted at orifice, 12–21 mm diam., rounded abaxially; stamens included or longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen sacs navicular, 1.6–2.4 mm, sutures smooth; staminode 25–30 mm, exserted, 0.9–1.1 mm diam., tip strongly recurved to coiled, distal 8–10 mm, and especially tip, moderately to densely lanate, hairs yellow, to 3.5 mm, proximal 6–8 mm glandular-puberulent; style 19–23 mm, glabrous or sparsely glandular-pubescent proximally. |
4–6 × 3–4 mm, glabrous. |
11–16 × 6–8 mm, sparsely glandular-pubescent distally. |
= 16. |
|
|
|
|
|
Flowering May–Aug. |
|
Gravelly slopes, cliffs, rocky clearings, roadcuts in coniferous forests. |
|
800–2800 m. (2600–9200 ft.) |
|
ID; MT; AB; BC
|
AZ; CA; CO; ID; NM; NV; UT; WA; WY
|
Penstemon caelestinus has been treated as a synonym of P. albertinus (D. D. Keck 1945; A. Cronquist 1959). D. V. Clark (1971) grew these and related species in transplant gardens and considered P. caelestinus to be distinct based on flower color, leaf shape and color, phenology, and habitat. A broad zone of hybridization between P. albertinus and P. wilcoxii exists in east-central Idaho and western Montana, especially in habitats disturbed by humans (Clark). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). Penstemon ×bryantiae D. D. Keck, a hybrid between P. palmeri and P. spectabilis, was described based on plants that volunteered in the Rancho Santa Ana Botanic Garden in 1935 (D. D. Keck 1937b). Penstemon ×mirus A. Nelson, a putative hybrid between P. eatonii and P. palmeri, has been reported from Arizona (A. Nelson 1938). A poultice prepared from Penstemon palmeri is applied to snakebites by the Kayenta Navajo of northeastern Arizona (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
1. Corolla tubes 7–8 mm; distal cauline leaves sessile, sometimes connate-perfoliate. | var. macranthus |
1. Corolla tubes 4–6 mm; distal cauline leaves connate-perfoliate. | → 2 |
2. Peduncles, pedicels, and calyx lobes glandular-pubescent; pollen sacs 1.8–2.4 mm. | var. palmeri |
2. Peduncles, pedicels, and calyx lobes glabrous; pollen sacs 1.6–2(–2.2) mm. | var. eglandulosus |
|
FNA vol. 17, p. 190. |
FNA vol. 17, p. 252. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
|
|
P. caelestinus |
|
Greene: Leafl. Bot. Observ. Crit. 1: 167. (1906) — (as Pentstemon) |
A. Gray: Proc. Amer. Acad. Arts 7: 379. (1868) — (as Pentstemon) |
| |