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sand dune penstemon, sand-dune beardtongue, sharp-leaf penstemon

Palmer's beardtongue, Palmer's penstemon, scented beardtongue, scented penstemon

Habit Herbs.
Stems

ascending to erect, (9–)20–60 cm, glabrous or ± scabrous, especially distally.

ascending to erect, 43–140 cm, glaucous.

Leaves

basal and cauline, glabrous, sometimes scabrous;

basal and proximal cauline (14–)30–95 × 4–15(–24) mm, blade oblanceolate to elliptic, base tapered, apex obtuse to acute or mucronate;

cauline (2 or)3–6(or 7) pairs, sessile, 18–60(–85) × (4–)8–28 mm, blade ovate to lanceolate, base tapered to cordate-clasping, apex acute to acuminate.

basal and proximal cauline (28–)52–120(–145) × (5–)25–50(–60) mm, blade ovate, base tapered, margins coarsely dentate, apex obtuse to acute;

cauline 5–8 pairs, sessile or connate-perfoliate, 25–130 × 7–45 mm, blade ovate to triangular, base of distals auriculate-clasping or connate-perfoliate, margins coarsely dentate, distals sometimes entire, apex acute.

Thyrses

continuous or interrupted, cylindric, 6–25(–30) cm, axis glabrous, rarely scabrous or glutinous, verticillasters (5–)7–15, cymes (2–)4–8(–12)-flowered;

proximal bracts ovate to lanceolate, 14–60 × 9–30 mm, margins entire, apex obtuse to acuminate or caudate;

peduncles and pedicels glabrous or scabrous, sometimes also glutinous, peduncles to 4 mm.

interrupted, rarely continuous, secund, 7–63 cm, axis glabrous or glandular-pubescent, verticillasters (4–)7–17, cymes (1 or)2–5-flowered;

proximal bracts ovate to lanceolate, 8–40 × 4–38 mm;

peduncles and pedicels ascending to erect, glabrous or glandular-pubescent.

Flowers

calyx lobes lanceolate, (3.2–)4.5–10 × 1.2–3.2(–3.8) mm, margins entire or erose, herbaceous or narrowly scarious, glabrous or scabrous proximally and, sometimes, along margins, sometimes also glutinous;

corolla blue to violet or purple, with reddish purple nectar guides, tubular-funnelform, 11–20 mm, glabrous externally, glabrous internally, tube 5–9 mm, throat gradually inflated, 4–6 mm diam., rounded abaxially;

stamens included or longer pair reaching orifice to slightly exserted, pollen sacs opposite, navicular, 0.7–1.3(–1.5) mm, sutures papillate;

staminode 6–12 mm, reaching orifice, 0.6–1 mm diam., tip recurved, distal 1.5–3.5 mm densely pilose, hairs golden yellow, to 0.6 mm, hairs on proximal part of staminode mostly along margins;

style 8–15 mm.

calyx lobes ovate, (3.4–)4.2–6(–7.5) × 1.9–3.6 mm, glabrous or glandular-pubescent;

corolla pale pink to white or pinkish lavender, with reddish purple nectar guides, strongly bilabiate, ventricose-ampliate, 25–38 mm, glandular-pubescent externally, glandular-pubescent internally, sometimes also sparsely white-lanate abaxially, tube 4–8 mm, length 1–1.4 times calyx lobes, throat abruptly inflated, constricted at orifice, 12–21 mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, filaments of shorter pair glandular-puberulent proximally, pollen sacs navicular, 1.6–2.4 mm, sutures smooth;

staminode 25–30 mm, exserted, 0.9–1.1 mm diam., tip strongly recurved to coiled, distal 8–10 mm, and especially tip, moderately to densely lanate, hairs yellow, to 3.5 mm, proximal 6–8 mm glandular-puberulent;

style 19–23 mm, glabrous or sparsely glandular-pubescent proximally.

Capsules

7–13 × 4–7 mm.

11–16 × 6–8 mm, sparsely glandular-pubescent distally.

Penstemon acuminatus

Penstemon palmeri

Distribution
from FNA
ID; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; NM; NV; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Penstemon acuminatus includes the northern var. acuminatus, which has denser thyrses than the southern var. latebracteatus. Herbage of var. acuminatus is usually scabrous; herbage of var. latebracteatus is rarely scabrous.

The Blackfoot tribe of the Central Rocky Mountains use Penstemon acuminatus as an analgesic, antiemetic, and gastrointestinal aid (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Penstemon ×bryantiae D. D. Keck, a hybrid between P. palmeri and P. spectabilis, was described based on plants that volunteered in the Rancho Santa Ana Botanic Garden in 1935 (D. D. Keck 1937b). Penstemon ×mirus A. Nelson, a putative hybrid between P. eatonii and P. palmeri, has been reported from Arizona (A. Nelson 1938).

A poultice prepared from Penstemon palmeri is applied to snakebites by the Kayenta Navajo of northeastern Arizona (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles 11–15 mm; corollas 14–20 mm; pollen sacs 0.8–1.3(–1.5) mm; calyx lobes 5–10 mm.
var. acuminatus
1. Styles 8–11 mm; corollas 11–15 mm; pollen sacs 0.7–1 mm; calyx lobes (3.2–)4.5–7 mm.
var. latebracteatus
1. Corolla tubes 7–8 mm; distal cauline leaves sessile, sometimes connate-perfoliate.
var. macranthus
1. Corolla tubes 4–6 mm; distal cauline leaves connate-perfoliate.
→ 2
2. Peduncles, pedicels, and calyx lobes glandular-pubescent; pollen sacs 1.8–2.4 mm.
var. palmeri
2. Peduncles, pedicels, and calyx lobes glabrous; pollen sacs 1.6–2(–2.2) mm.
var. eglandulosus
Source FNA vol. 17, p. 113. FNA vol. 17, p. 252.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei Plantaginaceae > Penstemon > subg. Penstemon > sect. Spectabiles
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. acuminatus var. acuminatus, P. acuminatus var. latebracteatus
P. palmeri var. eglandulosus, P. palmeri var. macranthus, P. palmeri var. palmeri
Name authority Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1285. (1829) — (as Pentstemon acuminatum) A. Gray: Proc. Amer. Acad. Arts 7: 379. (1868) — (as Pentstemon)
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