sand dune penstemon, sand-dune beardtongue, sharp-leaf penstemon
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cup-leaf beardtongue, scarlet beardtongue
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ascending to erect, (9–)20–60 cm, glabrous or ± scabrous, especially distally. |
ascending to erect, 50–100(–150) cm, glabrous. |
basal and cauline, glabrous, sometimes scabrous; basal and proximal cauline (14–)30–95 × 4–15(–24) mm, blade oblanceolate to elliptic, base tapered, apex obtuse to acute or mucronate; cauline (2 or)3–6(or 7) pairs, sessile, 18–60(–85) × (4–)8–28 mm, blade ovate to lanceolate, base tapered to cordate-clasping, apex acute to acuminate. |
basal and cauline, glabrous; basal and proximal cauline 40–100 × (8–)14–30 mm, blade spatulate to ovate, base tapered, apex rounded to obtuse, rarely retuse; cauline 5–9 pairs, sessile, 12–110 × 16–70 mm, blade oblong to ovate, base clasping to connate-perfoliate, apex obtuse to acute. |
continuous or interrupted, cylindric, 6–25(–30) cm, axis glabrous, rarely scabrous or glutinous, verticillasters (5–)7–15, cymes (2–)4–8(–12)-flowered; proximal bracts ovate to lanceolate, 14–60 × 9–30 mm, margins entire, apex obtuse to acuminate or caudate; peduncles and pedicels glabrous or scabrous, sometimes also glutinous, peduncles to 4 mm. |
interrupted, cylindric, 15–50 cm, axis glabrous, verticillasters 6–10(–16), cymes 1–5-flowered; proximal bracts ovate, 9–45 × 14–40 mm; peduncles and pedicels glabrous. |
calyx lobes lanceolate, (3.2–)4.5–10 × 1.2–3.2(–3.8) mm, margins entire or erose, herbaceous or narrowly scarious, glabrous or scabrous proximally and, sometimes, along margins, sometimes also glutinous; corolla blue to violet or purple, with reddish purple nectar guides, tubular-funnelform, 11–20 mm, glabrous externally, glabrous internally, tube 5–9 mm, throat gradually inflated, 4–6 mm diam., rounded abaxially; stamens included or longer pair reaching orifice to slightly exserted, pollen sacs opposite, navicular, 0.7–1.3(–1.5) mm, sutures papillate; staminode 6–12 mm, reaching orifice, 0.6–1 mm diam., tip recurved, distal 1.5–3.5 mm densely pilose, hairs golden yellow, to 0.6 mm, hairs on proximal part of staminode mostly along margins; style 8–15 mm. |
calyx lobes lanceolate, 4.5–7 × 1.8–2.9 mm, margins entire, narrowly scarious, glabrous; corolla red, without nectar guides, tubular-funnelform, 32–36 mm, glabrous externally, glabrous internally, tube 8–11 mm, throat abruptly inflated, 9–12 mm diam., rounded abaxially; stamens exserted, pollen sacs opposite, explanate, 1.3–1.7 mm, sutures smooth; staminode 18–20 mm, exserted, 0.2–0.3 mm diam., tip straight to recurved, glabrous; style 24–31 mm. |
7–13 × 4–7 mm. |
9–13 × 6–10 mm. |
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Flowering Mar–Jun. |
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Sandy soils, deciduous and pine woodlands, sandhill prairies. |
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10–200 m. (0–700 ft.) |
ID; NV; OR; WA
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AR; LA; OK; TX
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Varieties 2 (2 in the flora). Penstemon acuminatus includes the northern var. acuminatus, which has denser thyrses than the southern var. latebracteatus. Herbage of var. acuminatus is usually scabrous; herbage of var. latebracteatus is rarely scabrous. The Blackfoot tribe of the Central Rocky Mountains use Penstemon acuminatus as an analgesic, antiemetic, and gastrointestinal aid (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
F. S. Crosswhite and C. D. Crosswhite (1981) hypothesized that Penstemon murrayanus was derived from P. grandiflorus, with which it shares many morphologic features. The two species have been crossed artificially, yielding hybrids that once were commercially popular (G. Viehmeyer 1958; R. Nold 1999). Molecular data support the sister relationship of P. murrayanus and P. grandiflorus (C. A. Wessinger et al. 2016). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Styles 11–15 mm; corollas 14–20 mm; pollen sacs 0.8–1.3(–1.5) mm; calyx lobes 5–10 mm. | var. acuminatus |
1. Styles 8–11 mm; corollas 11–15 mm; pollen sacs 0.7–1 mm; calyx lobes (3.2–)4.5–7 mm. | var. latebracteatus |
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FNA vol. 17, p. 113. |
FNA vol. 17, p. 121. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
P. abietinus, P. absarokensis, P. acaulis, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1285. (1829) — (as Pentstemon acuminatum) |
Hooker: Bot. Mag. 63: plate 3472. (1836) — (as Pentstemon) |
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