sand dune penstemon, sand-dune beardtongue, sharp-leaf penstemon
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Canaan Mountain beardtongue, sandloving penstemon
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ascending to erect, (9–)20–60 cm, glabrous or ± scabrous, especially distally. |
decumbent to ascending, 5–26 cm, usually ± fistulose, glutinous and covered with sand, not glaucous. |
basal and cauline, glabrous, sometimes scabrous; basal and proximal cauline (14–)30–95 × 4–15(–24) mm, blade oblanceolate to elliptic, base tapered, apex obtuse to acute or mucronate; cauline (2 or)3–6(or 7) pairs, sessile, 18–60(–85) × (4–)8–28 mm, blade ovate to lanceolate, base tapered to cordate-clasping, apex acute to acuminate. |
basal and cauline, not leathery, glutinous and covered with sand, not glaucous; basal and proximal cauline 18–80 × 4–15 mm, blade oblanceolate, base tapered, margins entire, crisped, apex obtuse to acute; cauline 2–5 pairs, sessile, 15–75 × 2–8 mm, blade oblanceolate to lanceolate, base tapered to truncate, margins crisped, apex rounded or obtuse to acute. |
continuous or interrupted, cylindric, 6–25(–30) cm, axis glabrous, rarely scabrous or glutinous, verticillasters (5–)7–15, cymes (2–)4–8(–12)-flowered; proximal bracts ovate to lanceolate, 14–60 × 9–30 mm, margins entire, apex obtuse to acuminate or caudate; peduncles and pedicels glabrous or scabrous, sometimes also glutinous, peduncles to 4 mm. |
continuous, cylindric, 4–15 cm, axis glutinous and covered with sand, verticillasters 4–10, cymes 1–6-flowered, 2 per node; proximal bracts lanceolate, 14–45 × 2–5(–8) mm; peduncles and pedicels glutinous and covered with sand. |
calyx lobes lanceolate, (3.2–)4.5–10 × 1.2–3.2(–3.8) mm, margins entire or erose, herbaceous or narrowly scarious, glabrous or scabrous proximally and, sometimes, along margins, sometimes also glutinous; corolla blue to violet or purple, with reddish purple nectar guides, tubular-funnelform, 11–20 mm, glabrous externally, glabrous internally, tube 5–9 mm, throat gradually inflated, 4–6 mm diam., rounded abaxially; stamens included or longer pair reaching orifice to slightly exserted, pollen sacs opposite, navicular, 0.7–1.3(–1.5) mm, sutures papillate; staminode 6–12 mm, reaching orifice, 0.6–1 mm diam., tip recurved, distal 1.5–3.5 mm densely pilose, hairs golden yellow, to 0.6 mm, hairs on proximal part of staminode mostly along margins; style 8–15 mm. |
calyx lobes lanceolate, 5–6.5 × 1–2 mm, glutinous and covered with sand; corolla lavender to blue-lavender, with violet nectar guides, weakly ventricose, 14–17 mm, glabrous externally, usually glutinous and covered with sand distally, especially on lobes, glabrous internally, tube 5–6 mm, throat gradually inflated, not constricted at orifice, 5–6 mm diam., slightly 2-ridged abaxially; stamens: longer pair reaching orifice, pollen sacs opposite, navicular, 0.9–1.4 mm, dehiscing completely or incompletely, connective splitting or not, sides glabrous, sutures papillate; staminode: 8–9 mm, included, 0.3–0.5 mm diam., tip straight, distal 4–5 mm sparsely papillate, papillae violet, to 0.2 mm; style 10–12 mm. |
7–13 × 4–7 mm. |
3.5–6.5 mm. |
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Flowering May–Jun. |
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Sand dunes, ponderosa pine forests, mixed shrublands. |
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1800–2200 m. (5900–7200 ft.) |
ID; NV; OR; WA
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UT |
Varieties 2 (2 in the flora). Penstemon acuminatus includes the northern var. acuminatus, which has denser thyrses than the southern var. latebracteatus. Herbage of var. acuminatus is usually scabrous; herbage of var. latebracteatus is rarely scabrous. The Blackfoot tribe of the Central Rocky Mountains use Penstemon acuminatus as an analgesic, antiemetic, and gastrointestinal aid (D. E. Moerman 1998). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon ammophilus is known from Garfield, Kane, and Washington counties. Populations occur on aeolian sand, derived from the Navajo Sandstone. The fistulose stems of P. ammophilus are unique in Penstemon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Styles 11–15 mm; corollas 14–20 mm; pollen sacs 0.8–1.3(–1.5) mm; calyx lobes 5–10 mm. | var. acuminatus |
1. Styles 8–11 mm; corollas 11–15 mm; pollen sacs 0.7–1 mm; calyx lobes (3.2–)4.5–7 mm. | var. latebracteatus |
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FNA vol. 17, p. 113. |
FNA vol. 17, p. 161. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Coerulei |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
P. abietinus, P. absarokensis, P. acaulis, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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Douglas ex Lindley: Edwards’s Bot. Reg. 15: plate 1285. (1829) — (as Pentstemon acuminatum) |
N. H. Holmgren & L. M. Shultz: Brittonia 34: 381, fig. 1. (1982) |
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