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pearl millet, yellow bristlegrass

fountain grass

Habit Plants annual. Plants annual or perennial; habit various.
Culms

50-300 cm, erect, branching;

nodes glabrous.

3-800 cm, not woody, sometimes branching above the base;

internodes solid or hollow.

Sheaths

glabrous or pubescent, with or without ciliate margins;

ligules 2-5 mm;

blades 15-100 cm long, 7-70 mm wide, flat, glabrous or pubescent.

Panicles

terminal, 4-200 cm long, 2-70 mm wide, fully exerted from the sheaths, erect;

rachises terete, densely pubescent.

Inflorescences

spicate panicles with highly reduced branches termed fascicles;

panicles 1-many per plant, terminal on the culms or on both the culms and the secondary branches, or terminal and axillary, or only axillary, usually completely exposed at maturity;

rachises usually terete, with (1)5-many fascicles;

fascicle axes 0.2-7.5(28) mm, with (1)3-130+ bristles and 1-12 spikelets.

Spikelets

3-7 mm;

pedicels 0.6-1.8 mm;

lower glumes absent or to 1.5 mm, veinless;

upper glumes 0.5-3.5 mm, 3-5-veined;

lower florets staminate or sterile;

lower lemmas 1.5-6 mm, glabrous, 3-7-veined, margins ciliate;

lower paleas vestigial or fully developed, margins ciliate;

anthers 2.2-2.5 mm, penicillate;

upper florets coriaceous, shiny;

upper lemmas 4.3-7 mm, 5-7(9)-veined, margins ciliate;

upper paleas 3.4-3.9 mm, pubescent, at least near the base, margins ciliate;

anthers 2-2.2 mm, penicillate.

with 2 florets;

lower glumes absent or present, 0-5-veined;

upper glumes longer, 0-11-veined;

lower florets sterile or staminate;

lower lemmas usually as long as the spikelets, membranous, 3-15-veined, margins usually glabrous;

lower paleas present or absent;

upper lemmas membranous to coriaceous, 5-12-veined;

upper paleas shorter than the lemmas but similar in texture;

lodicules 0 or 2, glabrous;

anthers 3, if present, x = 5,7, 8,9 (usually 9).

Caryopses

2-5.5 mm long, 1.6-3.2 mm wide, protruding from the lemma and palea at maturity.

Fascicles

33-160 per cm;

fascicle axes 1-28 mm, persistent, with 1-9 spikelets;

outer bristles 44-131, 0.5-6 mm;

inner bristles 6-19, 4-6 mm, plumose;

primary bristles 5.5-6.3, ciliate, sometimes noticeably longer than the other bristles.

Ligules

membranous and ciliate, or of hairs, rarely completely membranous;

blades sometimes pseudopetiolate.

Bristles

free or fused at the base, disarticulating with the spikelets at maturity; of 3 kinds, outer, inner, and primary, in some species with all 3 kinds present below each spikelet, in others 1 or more kinds missing from some or all of the spikelets;

outer (lower) bristles antrorsely scabrous, terete;

inner (upper) bristles antrorsely scabrous or long-ciliate, usually flatter and wider than the outer bristles;

primary (terminal) bristles located immediately below the spikelets, solitary, antrorsely scabrous or long-ciliate, often longer than the other bristles associated with the spikelet;

disarticulation usually at the base of the fascicles, sometimes also beneath the upper florets.

2n

= 14.

Pennisetum glaucum

Pennisetum

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; PR
[WildflowerSearch map]
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; PR; Virgin Islands
[BONAP county map]
Discussion

Pennisetum glaucum, a native of Asia, is cultivated in the United States for grain, forage, and birdseed. It is the most drought tolerant of the tropical cereal crops. Under favorable conditions, 10,000-30,000+ fascicles may be produced. In the Flora region, it is used for soil stabilization, partly because it seldom persists for more than 1-2 years.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pennisetum has 80-130 species, most of which grow in the tropics and subtropics, and occupy a wide range of habitats. Twenty-five species are native to the Western Hemisphere, but none to the Flora region. Most of the species treated here are cultivated for food, forage, or as ornamental plants. Many species, including several cultivated species, are weedy. Four are classified as noxious weeds by the U.S. Department of Agriculture. Records known to be based on cultivated plants are not included in the distribution maps but, in many cases, it is not possible to determine whether a record is based on a cultivated plant or an escape.

The placement of the boundary between Pennisetum and Cenchrus is contentious. As treated here, Pennisetum has antrorsely scabrous bristles that are not spiny, fascicle axes that terminate in a bristle, and chromosome base numbers of 5, 7, 8, and 9. Cenchrus has retrorsely (rarely antrorsely) scabrous, spiny bristles, fascicle axes that are terminated by a spikelet, and a chromosome base number of 17 (Wipff 2001). In both genera, the bristles are reduced branches (Goebel 1882; Sohns 1955).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants stoloniferous; panicles axillary, partially or wholly hidden in the leaf sheaths at maturity, the rachises flattened in cross section, with 1-6 fascicles; spikelets 10-22 mm long, bristles mostly shorter than the spikelet
P. clandestinum
1. Plants not stoloniferous; panicles terminal or terminal and axillary, fully exserted at maturity, the rachises terete, with 10-many fascicles; spikelets 2.5-12 mm long, the majority of the bristles as long as or longer than the spikelets.
→ 2
2. Fascicles with only 1 bristle and 1 spikelet
P. petiolare
2. Fascicles with 6 or more bristles and 1-12 spikelets.
→ 3
3. Most or all bristles scabrous, the primary bristles sometimes sparsely and inconspicuously long-ciliate.
→ 4
4. Primary bristles not noticeably longer than all the other bristles in the fascicles.
→ 5
5. Terminal panicle erect; fascicles with a stipelike base 1.5-5.6 mm long
P. alopecuroides
5. Terminal panicle drooping; fascicles subsessile, the bases 0.4-0.7 mm long.
→ 6
6. Plants green; most of the bristles only slightly longer than the spikelets; upper glumes (7)9-veined, about as long as the spikelets
P. nervosum
6. Plants purplish; bristles at least twice as long as the spikelets; upper glumes 1-3-veined, usually about 1/2 as long as the spikelets
P. macrostachys
4. Primary bristles noticeably longer than all the other bristles in the fascicles.
→ 7
7. Panicles dense; rachises with 21-40 fascicles per cm.
→ 8
8. Rachises pubescent; bristles yellow or purple; leaf blades (4)12-40 mm wide; paleas of lower florets present
P. purpureum
8. Rachises scabrous; bristles white to stramineous; leaf blades 4-12 mm wide; paleas of lower florets absent
P. macrourum
7. Panicles less dense; rachises with 5-16 fascicles per cm.
→ 9
9. Panicles drooping, terminal and axillary; leaf blades 19-45 mm wide
P. latifolium
9. Panicles erect, all terminal; leaf blades 2-12 mm wide.
→ 10
10. Plants not rhizomatous; lower part of rachises pubescent
P. alopecuroides
10. Plants rhizomatous; lower part of rachises scabrous
P. flaccidum
3. Bristles, at least the primary bristles, conspicuously long-ciliate.
→ 11
11. Spikelets 9-12 mm long
P. villosum
11. Spikelets 2.5-7 mm long.
→ 12
12. Fascicles not disarticulating from the rachises; panicles 4-200 cm long; upper lemmas with pubescent margins; caryopses protruding from the florets at maturity
P. glaucum
12. Fascicles disarticulating from the rachises at maturity; panicles 2-37.5 cm long; upper lemmas with glabrous margins; caryopses concealed by the lemmas and paleas at maturity.
→ 13
13. Upper florets readily disarticulating at maturity; upper lemmas smooth and shiny, conspicuously different in texture from the lower lemmas.
→ 14
14. Fascicles with 6-14 long-ciliate inner bristles and 13-30 scabrous outer bristles; fascicle axes 0.2-0.5 mm long; spikelets sessile
P. polystachion
14. Fascicles with 40-90 long-ciliate inner bristles and 10-20 scabrous outer bristles; fascicle axes 1.5-2.5 mm long; spikelets pedicellate, the pedicels 1-3.5 mm long
P. pedicellatum
13. Upper florets not disarticulating at maturity; lower and upper lemmas similar in texture.
→ 15
15. Lower portion of the rachises glabrous, sometimes scabrous.
→ 16
16. Inner bristles neither grooved nor fused, even at the base; spikelets 5.2-6.7 mm long, pedicellate, the pedicels 0.1-0.5 mm long
P. flaccidum
16. Inner bristles grooved and fused, at least at the base; spikelets 2.5-5.6 mm long, sessile.
→ 17
17. Inner bristles fused for up to 1/4 their length; many outer bristles exceeding the spikelets; terminal bristles 10.5-23 mm, noticeably longer than the other bristles in the fascicles
P. ciliare
17. Inner bristles fused for 1/3 – 1/2 their length; outer bristles not exceeding the spikelets; terminal bristles 2.9-6.5 mm, usually not noticeably longer than other bristles in the fascicles
P. setigerum
15. Lower portion of the rachises pubescent.
→ 18
18. Plants 200-800 cm tall; midculm leaves (4)12-40 mm wide; panicles golden-yellow or dark purple; rachises with 30-40 fascicles per cm
P. purpureum
18. Plants 50-200 cm tall; midculm leaves 2-11 mm wide; panicles white, burgundy, light purple, or pink; rachises with 5-17 fascicles per cm.
→ 19
19. Midculm leaves 2-3.5 mm wide, convolute or folded, green, the midvein noticeably thickened; lower florets of the spikelets usually sterile, sometimes staminate
P. setaceum
19. Midculm leaves 3-11 mm wide, flat, green or burgundy, the midvein not noticeably thickened; lower florets of the spikelets staminate.
→ 20
20. Plants shortly rhizomatous; nodes pubescent; panicles erect to slightly arching, white or purple-tinged; leaves green; ligules 1-1.7 mm long; fascicles with 0-24 terete, scabrous outer bristles
P. orientale
20. Plants not rhizomatous; nodes glabrous; panicles conspicuously drooping, burgundy (rarely whitish-green); leaves burgundy (rarely green); ligules 0.5-0.8 mm long; fascicles with 43-68 terete, scabrous outer bristles
P. advena
Source FNA vol. 25. FNA vol. 25, p. 515. Author: J.K. Wipff;.
Parent taxa Poaceae > subfam. Panicoideae > tribe Paniceae > Pennisetum Poaceae > subfam. Panicoideae > tribe Paniceae
Sibling taxa
P. advena, P. alopecuroides, P. ciliare, P. clandestinum, P. flaccidum, P. latifolium, P. macrostachys, P. macrourum, P. nervosum, P. orientale, P. pedicellatum, P. petiolare, P. polystachion, P. purpureum, P. setaceum, P. setigerum, P. villosum
Subordinate taxa
P. advena, P. alopecuroides, P. ciliare, P. clandestinum, P. flaccidum, P. glaucum, P. latifolium, P. macrostachys, P. macrourum, P. nervosum, P. orientale, P. pedicellatum, P. petiolare, P. polystachion, P. purpureum, P. setaceum, P. setigerum, P. villosum
Name authority (L.) R. Br. Rich.
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